CILIATE FIBRILLAR SYSTEMS 199 



"completely to demonstrate the great pharyngeal complex," which is 

 an integral part of this system. 



Lund was able to differentiate between these "different aggregations 

 of structures" by means of the silverline technique (Klein's and von 

 Gelei and Horvath's) on the one hand and, on the other, by the use of 

 iron-haematoxylin and Mallory's stain. 



The former method demonstrated von Gelei's "Stiitzgitter System" 

 and Klein's "Indirekt verbindung System," i.e., their "outer fibrillar 

 complex" noted above. It also revealed essentially their "inner fibrillar 

 complex," also as noted above. 



But, according to Lund, these are separate and distinct "aggregates." 

 The "outer fibrillar complex" is not subpellicular, as both von Gelei 

 and Klein maintain, but represents rather the sculptural polygonal pat- 

 tern of the pellicle itself. A similar interpretation was made by Brown 

 (1930). 



The "inner fibrillar complex" of von Gelei and of Klein comprises the 

 basal granules, their connecting longitudinal body fibrils, transverse 

 fibrils connecting the longitudinal fibrils, and others which include the 

 "radial fibrils." These last mentioned "originate as longitudinal fibrils 

 in the cytopharynx and oesophagus, spread radially out from the oral 

 opening over the body surface and terminate a short way from the cyto- 

 stome." 



This inner fibrillar complex may be clearly demonstrated by the silver- 

 line techniques, especially by the wet method. There is, however, a 

 portion of the fibrillar complex within the cytopharynx and cytoesopha- 

 gus which is not wholly demonstrable by these techniques. This was well 

 differentiated by the iron-haematoxylin and Mallory's methods, and 

 described as "seven major parts, namely: (1) the pharyngo-esophageal 

 network, (2) the neuromotorium, (3) the penniculus, (4) the oesopha- 

 geal process, (5) the paraesophageal fibrils, (6) the posterior neuromotor 

 chain, and (7) postesophageal fibrils." 



For the descriptive details of this very elaborate complex of fibrils 

 and associated parts, obviously the original account must be read. It is 

 evident, however, from this brief review of the results of these several 

 workers on the fibrillar system of Paramecium that a number of discrep- 

 ancies need to be cleared up and perhaps further structural analysis of 



