228 CILIATE FIBRILLAR SYSTEMS 



substance, the "fibrillare Komponente" (1928, p. 255). By virtue of its 

 integral relation with the basal apparatus of the cilia and with the tricho- 

 cysts, he attributed to his silverline system also a specifically conductive, 

 coordinative function. 



Chatton and Lwoff's (1935-36) criticisms of Klein's interpretations 

 bear mainly upon the structural relations of the fibril to which are ad- 

 joined, always on its left side, the basal granules. The former authors 

 think Klein's silverline system is quite separate and distinct from their 

 infraciliature. It appears to the writer, however, that this discrepancy 

 might be completely resolved by identifying Chatton and Lwoff's infra- 

 ciliature with the longitudinal fibrils and basal granules of Klein and 

 von Gelei's inner fibrillar complex. Further critical investigation would 

 be needed, of course, to establish such identity. 



Fibrillar Systems of Other Ciliates 

 a. holotricha 



Ancistruma (Kidder, 1933). — The fibrils in Ancistruma mytili and 

 A. isseli are of three types — the longitudinal fibrils of the ciliary rows, 

 the irregularly distributed transverse fibrils, and the net complex of fine 

 fibrils in the peristomal region. The latter fibrils seem to connect di- 

 rectly with the basal bodies of the peristomal cilia, and they are coarser 

 and less numerous in A. isseli than in A. mytili. The longitudinal fibrils 

 in A. mytili are continuous around the posterior end, but in A. isseli 

 they center about a posterior suture. 



A number of fine fibrils connect the inner dorsal row of peristomal 

 cilia to the outer dorsal row. These seem to be distinct from the net 

 complex and probably are in the nature of concentrated transverse 

 fibrils. In A. isseli such fibrils are absent, and instead the fibril of the 

 outer ciliary row joins that of the inner dorsal row. 



Fibrils resembling the interstrial fibrils of Boveria (Pickard, 1927) 

 are sometimes seen in A. mytili, but these may represent a deep-lying 

 network of the same type regularly seen in the peristomal region. 



Methods 



Fixatives: Schaudinn, sublimate-acetic, Bouin's, Zenker's, Champy's. 

 Stains: Heidenhain's and Delafield's haematoxylin, crystal violet-sulph- 

 alizarinate (Benda's). 



