230 CILIATE FIBRILLAR SYSTEMS 



fibrils of the neuromotor system stained bright red by Mallory's method. 

 When dislodged along with the basket by means of microdissection, the 

 motorium was "a refringent body." In stained sections, however, it ap- 

 peared granular. 



"Fans of fibrils" join the motorium at both its anterior and posterior 

 ends. Those at the anterior end mark the convergence of longitudinal 

 fibrils (Fig. 81 A) of the body cilia, of ventral fibrils from the posterior 

 region of the body and of fibrils from the mouth. The circular myonemes 

 of the mouth are traversed by many fine fibrils that continue on toward 

 the "railroad track." A fibrillar fan from the posterior end of the moto- 

 rium continues as the dorsal fibers to the "railroad track," which, in turn, 

 show a very complex system of fibrils associated with its trichites. 



The basal bodies of the cilia are connected not only by the longitudinal 

 fibrils but also by cross fibrils (Fig. 8 IB). 



By methods of microdissection it was shown that (1) after destruc- 

 tion of the motorium "there is a marked disturbance in the action of the 

 cilia, in no way comparable to the disturbance of the cilia if other parts 

 of the body are injured"; (2) the cilia may still exhibit a wave-like 

 motion, but they do not reverse after the motorium is destroyed. This 

 seemed to suggest a coordinating function for the fibrillar system of this 

 Chlamydodon, whose inconspicuous motor organelles are not favorable 

 for an experimental study of modifications of their coordinated activity. 



Methods 



Fixatives: Schaudinn's, Bouin's, and strong Flemming's. 



Stains: Iron-haematoxylin, Mallory's (after Zenker's or picromercuric 



fixation). 

 Microdissection. 



Conchophthirus mytili De Morgan (Kidder, 1933). — The peripheral 

 longitudinal fibrils linking the ciliary basal bodies originate from a 

 transverse fibril in the anterior ventral region. These almost parallel rows 

 of fibrils pass around the posterior end of the organism uninterrupted, 

 continue over the anterior end, and again return to the transverse fibril 

 of the ventral surface. Each basal body is furnished with a ciliary rootlet 

 extending toward the endoplasm. 



An elongate mass of homogeneous material, the motorium, follows the 

 posterior line of the cytopharynx and continues into the endoplasm as a 



