1920.] G. A. BOULENGER : Frogs of the Genus Rana. 93 



Foot .. .. 43 38 42 39 39 38 56 54 S6 55 54 48 48 



Third toe .. .. 24 20 24 21 2 1 20 29 28 30 30 28 27 26 



Fourth toe . . . . 36 32 35 36 32 3° 46 45 4^ 47 43 4' 40 



Fifth toe .. .. 27 23 27 25 25 21 34 35 34 34 32 31 30 



Habitat. Yunnan fu, Yunnan. Tj'pes in the British Museum. 



This species is very similar to R. andersonii, Blp,r., but readily 

 distinguished by the absence of discs to the fingers and toes. It 

 is truly a link between the subgenera Rana and Hylorana, and I 

 cannot doubt that a direct genetic relationship exists between 

 these two allied species ; this remark applies also to R. pleii- 

 raden and R. adenopleura, and points to the polyphyletic origin 

 of the subgenus Hylorana , which appears to be made up of 

 specialized forms leading to scansorial types all derived from the 

 Ranac typicae. 



43. Rana japonica. 



Rana femporaria {non I.inn.), Schleg., Fatin. Japan., Rept. p. log, pi. ii, 



fig. 2(1838). 

 Rana teniporaria, part., Middend., Stbir. Reis. II, p. 247, pi. xxvi, figs. 2 — 4 



(1853); Dum. at Bibr., Erp. Gen. VIII, p. 358 (1841). 

 Rana temporaria, var. japonica, Giinth., Cat. Batr. Sal. p. 17 (1858). 

 Rana silvatica, part., Hallow., Proc. Ac. Pliilad. i860, p. 500; Giinth., 



Rept. Brit.Ind. p. 409 (1864)1 Camerano, Atti. Ace. Torin. XIV, 1879, 



p. 876. 

 Rana middendorffii, Steenstr., Vid. Meddel. 1869, p. 18, fig. 

 Rana japonica^ Bouleng., Bull. Soc. Zooi. France, 1879, p. 190, and Cat. 



Batr. Ecaiid. p. 47 (1882) ; Boettg , Ber. Offenb. Ver. Nat. 1888, 



p. 96 ; Bouleng., Proc. Zool. Soc. 1891, p. 610; Boettg. Ber. Offenb. 



Ver. Nat. 1895, p. 102; Werner, Abli. Bay. Ak. XXII, 1904, p. 358; 



Stejneg., Herp.Jap. p. 107, fig (1907). 

 Rana japonica, var. ornativentris, Werner, t. c. p. 383. 

 Rana tsushimensis, Stejneg. op. cit. p. 116, fig. 



Vomerine teeth in oblique groups or short series between the 

 choanae or behind the level of their posterior borders. 



Head rather depressed, as long as broad or a little longer or 

 a little broader; snout rounded or more or less acuminate, more 

 or less projecting be3^ond the mouth, as long as or Httle longer 

 than the eye; canthus rostralis distinct; loreal region oblique, 

 feebly concave; nostril equidistant from the eye and from the tip 

 of the snout, or a little nearer the latter ; distance between the 

 nostrils equal to or a little greater than the interorbital width, 

 which equals I to t, rarely once, that of the upper eyelid and is 

 usually less than the diameter of the tympanum; tympanum 

 very distinct, f to I the diameter of the eye and 2 to 3 times its 

 distance from the latter. 



Fingers moderate or rather long, obtuse, first as long as or 

 longer than the second, third longer than the snout ; subarticular 

 tubercles rather large, very prominent. 



Hind limb long, the tibio-tarsal articulation reaching the 

 nostril, the tip of the snout, or a little beyond, the heels strongly 

 overlapping when the limbs are folded at right angles to the body ; 

 tibia 4 to 51 times as long as broad, if to 21, times in length 



