56 THE NORTH AMERICAN SPECIES OF DROSOPHILA. 
The curious produced eyes of some of the males of Z. dispar caused 
Wiedemann to describe the species as a diopsine. They do not occur in 
the female of this species, or in either sex of the other species. Similar 
eyes are to be found in some undescribed Neotropical species of Chymomyza, 
close to C. procnemis. The present group is not close to Chymomyza in 
other respects. 
The two American species have both been bred from fungi. 
Dettopsomyia Lamb. 1914. Trans. Linn. Soc. London, 16, 349. 
Arista plumose; two orbitals, upper reclinate, lower proclinate; postverticals crossed; 
vibrisse and ocellars present; eyes pilose; carina large and broad; two dorsocentrals, plus 
two small presutural ones; acrostichal hairs in two rows; two humerals; one presutural; 
two pairs of scutellars, posterior ones crossed; two sternopleurals; costa reaches apex of 
fourth vein, twice broken, distal break deep, milichiid-like; costal index a little less than 1.0. 
There is a single described species, D. formosa Lamb, from the Seychelles. 
Camilla Haliday. 1838. Ann. Nat. Hist., 2, 188. 
Arista plumose; orbitals disposed as usual, middle one minute; postverticals medium 
size, convergent; vibrisse present; two notopleurals; mesopleure bristly; two dorso- 
centrals; no prescutellars; many acrostichal rows of hairs; presuturals long; two pairs of 
scutellars; costa twice broken; anal vein absent; anal cell open at apex; auxiliary vein 
rudimentary; no preapicals on third tibie. 
The type species is the European C. glabra Fallén. Seven species are 
described, as follows: 
Palearctic: C. glabra Fallén, Europe, Canary Islands; C. acutipennis Loew, Europe. 
Ethiopian: C. africana Bezzi, Kongo. 
Oriental: C. ceruleifrons de Meijere, Java; C. javana de Meijere, Java; C. pusilla de 
Meijere, Java; C. rugulosa de Meijere, Java. 
I have examined both European species. 
Stegana Meigen. 1830. Syst. Beschr., 6, 79. . 
Phortica Schiner. 1862. Wien. ent. Monatsschr., 6, 433. 
Amiota Loew. 1862. Berlin. ent. Zeit., 6. 
Hostegana Hendel. 1913. Deutsch. ent. Zeit., 390. 
Orthostegana Hendel. 1913. Deutsch, ent. Zeit. 
Arista plumose; three large orbitals, placed high up, uppermost one nearer to verticals 
than to lowest (proclinate) orbital; postverticals small; eyes nearly bare; vibrisse and 
ocellars present; one humeral; one presutural; two dorsocentrals; prescutellars present; 
two scutellar pairs; mesopleure bare; preapicals on all tibie; discal and second basal 
cells separated; costa reaches fourth vein; anal cell and anal vein present. 
I have ventured to unite the two long-recognized genera Stegana and 
Phortica. 'They are usually (e. g., by Schiner, by Williston, and by Olden- 
berg) stated to differ in that Stegana has wings bent down at the base and 
has the third and fourth veins strongly convergent, while in Phortica the 
wings are straight and the third and fourth veins are parallel or nearly so. 
But these two characters are not always associated. Hendel has described 
two new genera: Hostegana, with bent-down wings but first posterior cell 
not narrowed; Orthostegana, with wings straight but first posterior cell 
narrowed. ‘The narrowing of the first posterior cell is a character that shows 
variations from species to species, so that it is hard to know where to draw 
the line between ‘“‘narrowed” and “not narrowed.’ The bending down 
of the wings is still more elusive, at least in pinned material. I am con- 
vinced, from an examination of many specimens, including the types, that 
Phortica vittata Coquillett is based on specimens of typical Stegana in which 
the wings are straight. I have seen specimens in which one wing was 
straight, the other bent down. Specimens of Phortica humeralis Loew 
sometimes have the wings bent down. These two genera are perhaps dis- 
