SYSTEMATIC ACCOUNT. Lr 
tinct; in general appearance and in the shape of the head two types certainly 
exist, but the wing characters that have been used as the only means of 
separating them are evidently not valid. 
The type species of Stegana is S. coleoptrata Scopoli, a European species, 
designated as type by Westwood (1840, Intr., 2,153). The type of Phortica 
is the European Drosophila variegata Fallén, ‘which was the only species 
included in the genus originally. 
The group here considered as forming the genus Stegana includes nine 
described Palearctic species (Europe), five Neotropical (St. Vincent, Peru, 
Bolivia; I have seen the genus from Mexico and Central America), and 
twelve Oriental (Ceylon to Formosa and New Guinea). I have seen an 
Ethiopian specimen, collected by the American Museum Kongo Expedition. 
Our knowledge of the Nearctic species is in a very unsatisfactory condition, 
and I am not sure that the treatment given below will stand after more 
study. Six Nearctic species are recorded, but I am unable to recognize 
more than two in the large number of specimens that I have seen. 
Stegana coleoptrata Scopoli. 1763. Ent. Carniol., 338 (Musca). 
S. hypoleuca Meigen. 1830. Syst. Beschr., 6 (7). 
Phortica vittata Coquillett. 1901. Proc. U.S. Nat. Mus., 23. 
Loew recognized S. curvipennis Fallén (as S. nigra Meigen) from North 
America; I have seen a specimen so labeled in the Loew collection, from 
Pennsylvania. I am unable to see any good reason for separating this 
specimen from S. coleoptrata, though it appears from the accounts of Schiner 
and of Oldenberg that in Europe the two species are quite distinct. The 
European species seem to have been little understood in Loew’s time. It is 
now customary to identify the darker specimens as S. coleoptrata, the lighter 
ones as P. vittata; but intermediates occur; and I can find no other char- 
acter that will separate the supposed two species. 
Specimens examined: Mount Desert, Maine (C. W. Johnson); Mount 
Washington (Mrs. Slosson), Franconia (Mrs. Slosson), Bretton Woods 
(C. W. Johnson), New Hampshire; Brattleboro, Vermont (C. W. Johnson); 
Brookline, Fall River, Massachusetts (C. W. Johnson); Buttonwoods, 
Rhode Island (C. W. Johnson); Danbury, Winnipauk, Middletown, Roway- 
ton, Darien, Connecticut (C. W. Johnson); South Wales (M. C. Van- 
Duzee), Ithaca (S. W. Frost), New York; Delaware Water Gap (type of 
vittata), Clemonton (C. W. Johnson), Avalon (C. W. Johnson), New 
Jersey; Polk County, Wisconsin (C. F. Baker); Great Falls, Virginia (N. 
Banks); Daytona, Florida (C. W. Johnson). 
Recorded in Europe from England to Hungary. 
Stegana humeralis Loew. 1862. Centuria, 2, 93 (Amiota). 
A. leucostoma Loew. 1862. Centuria, 2, 94. 
(?) Drosophila alboguttata Wahlberg. 1838. K. Vet. Ak. Hand., 22. 
The types of Loew’s two species are quite distinct in color. S. humeralis 
is somewhat shining black in general color, while S. leucostoma is brown. 
I can find no other differences, and an extensive series of more recent 
material shows. graded color variations completely connecting these two 
extremes. Wahlberg’s species was recognized from the Nearctic by Loew. 
Two specimens so labeled in the Loew collection (from New Hampshire 
and New York, respectively), appear to me to belong to the same species 
as the type of S. humeralis. I am, however, unwilling to substitute Wahl- 
berg’s name without more study. 
Specimens examined: Waubamic, Parry Sound, Ontario (Aldrich coll. 1 
Franconia (Mrs. Slosson), Center Harbor (HEG: Dyar), Glen House (C. W 
