280 Records of the Indian Museum. [Vol. XV, 



metallica, Alex., Proc. V. S. Nat. Mus. XLIX, p. 178 (1916). 

 Java. 

 Alexander states in this paper that my Cylindrotoma quadricellulu 

 is a Stibadocera . 



Genera in CYLINDROTOMINI. 



An examination of the genera and species in this group reveals a 

 remarkable elasticity of characters and substantiates its intermediate 

 position between Tipulinae and Limnobiinae. 



This was quite evident to Osten Sacken who,^ in comparing the 

 European Cylindrotoma distinctissima with the North American ameri- 

 cana, also the European glabrata with the North American nodicornis, 

 wrote " The fact is that these species represent a gradation which baffles 

 every attempt at a generic arrangement." He retained Phalacrocera 

 tipulina and the European P. replicata in the same genus in spite of the 

 important difference that in the former the 1st vein ends in the 2nd 

 vein, with the marginal cross vein absent, whilst in replicata the 1st vein 

 ends in the costa with the marginal cross vein present. He kept all the 

 species known at that time in the three genera recognised by Schiner, 

 Cylindrotoma, Phalacrocera and Triogma " in order to avoid the estab- 

 lishment of a new genus for almost every species known, which would 

 probably necessitate a similai' process for every species to be discovered 

 hereafter." He notes that even the absence of the anterior cross vein 

 (when it is normally absent) is not always constant in the same species, 

 as out of twenty-one examples of Liogma nodicornis examined it was 

 absent in seventeen and present, though short, in four, so he retains 

 nodicornis and glabrata (in which latter the vein is present) in the same 

 genus Liogma. 



The characters of this group which exhibit such unusual variation 

 are : (1) the exact manner in which the auxiliary vein terminates, 

 with the presence or absence of the subcostal cross vein, or the presence 

 or otherwise of a short cross vein between the tip of the auxiliary vein 

 and the costa ; (2) the exact manner in which the 1st vein terminates, 

 with the presence or absence of what I call in my Fauna volume the 

 costal cross vein ; (3) the point at which the 2nd and 3rd veins diverge, 

 and, (4) the presence or absence of the anterior cross vein. Other 

 characters though variable are definite one way or the other, such as 

 the number (four or five) of posterior cells, the jDunctulate nature of the 

 thorax or otherwise, and some minor ones. All the tibiae are spurred 

 in the three species before me. 



The material before me, apart from literature, consists of a single 

 specimen of C. distinctissima (with one wing only), the three original 

 specimens of my C. 4-cellula,- and four ^ ^ and one $ of my new species 

 latefurcata. These exhibit the following comparative characters. 



1 Monog. N. Am. Tipulidae, p. 295. 



2 This is a Stibadocera as Alexander notes. I had not seen Eadorlein's peeper, nor an 

 earlier paper by Alexander on Neotropical Limnobiinae in which he doubts the Cylindro- 

 tomine character of the genus, though he admits its position here in his subsequent paper 

 on Javan TipuUdae. 



