192 Records of the Indian Museum. [VOr nae 
been described in Helix pomatia and other pulmonates of the 
same family (7, 8). 
Minute structure (pl. xxvii, figs. 27 and 28).—The tube is 
composed of the following coats from without inwards :— 
(1) A thin layer of connective tissue. 
(2) A layer of longitudinal muscle-fibres. 
(3) A layer of circular muscle-fibres. 
(4) A submucous coat of connective tissue (with a few 
muscle-fibres) raised into a number of longitudinal 
folds, generally ten, projecting into the lumen. 
(5) A single layer of columnar epithelium with numerous 
goblet cells (secreting cells). 
(2) The fenial sheath consists of two portions:—(1) A 
stout hollow spindle-shaped structure. giving attachment to the 
retractor muscle at its proximal end; (2) a stout tubular struc- 
ture arising from the distal end of the first portion and ending 
in the external aperture at the base of the right lower tentacle. 
Minute structure.—The tubular portion consists of alternate 
layers of longitudinal and transverse muscles arranged irregularly. 
The wall is thrown into large longitudinal folds, the core of which 
is formed mainly of longitudinal muscle fibres. The presence 
of these large folds shows the great extensibility of the penial 
sheath. The epithelium consists of a single layer of cubical (or 
short columnar) cells. 
(3) The penis is a more or less cylindrical structure iying 
in the proximal portion of the penial sheath when retracted. 
The penis is attached to the sheath at the proximal end and 
is traversed by a fine channel continuous with the lumen of 
the tube described above, and opening into the tip by a small 
aperture. 
(4) The refractor penis muscle is a short thick band extend- 
ing from the inner side of the body wall on the right, a little 
behind the female genital aperture, to the proximal end of the 
penial sheath and penis. 
(5) The Simrothian glands (pl. xxvi, figs. 19, 20a) are two 
tubular structures, one on each side, opening on the outer side of the 
lower tentacles, the right one being placed close to the male genital 
aperture. Each gland can be divided into four portions—(1) 
a long stout tubular portion coiled in various ways; (2) a short 
narrow portion, also tubular in structure, coiled closely ; (3) an 
elongated slightly curved conical portion continuous with the 
second one at the tapering end and ending in the next in a 
broad base; (4) a very short narrow tube with a small cylin- 
drical process from the outer side. 
The structure of the Simrothian glands in A. maximus and 
A. savasini resembles closely that of the present species except 
that there is no distinct fourth portion and the process arises 
from the proximal end of what corresponds to the third division 
