76 DEVELOPMENT OF THE FLOWER. 
parts must exist, unless it ean be ds that similar parts 
once formed, may be absorbed. 
The presence of the rudiment of a style, is in my opinion 
proof of there being an additional abortive carpellary leaf. 
'The other perfect carpella may be vascular, the rudiment 
may not only be evascular, but that part of the ovarium 
below it, which should be its lamina may also be evascular, - 
yet the proof would not be fatally shaken. I can easily 
conceive that processes may originate from an ovarium, but 
in these cases especial regard must be had to their situation, 
as for instance in Graminee, and if belonging to developed 
carpella, they will not be single, unless they grow from the 
dorsum. 
Penetration before the ovulum is reached, must only take 
place in a stigma in which the component parts are in a state 
of complete adhesion, or in which, as in Anagallis, the canal 
is stopped up by stigmatic tissue, 
This is seldom, if ever the case; in compound styles a 
central depression exists leading directly into the stigmatic 
canal, or this may have a distinct aperture; in simple styles 
the access to the canal of communication is generally very 
easy. In these cases what is the course of the £ogaux ? Do 
they pass over the stigmatic tissues into the canal, or through ; 
them? In such instances as Meyenia etc. ponettation can 
hardly be allowed. 
As a definition of value, in, putting one on one's guard 
against laxity of description of stigmata, I may cite Lindley's. 
*' Nothing is properly speaking stigma, except the secreting 
surface of the style." Although there is a want of precision 
in the term secreting, there is always a peculiar aspect of the 
stigmatic surface, by which it may be known from that of 
the style. 
Somewhat of a similar kind of misplacement of the stig- 
matic tissue occurs in certain Meliacez, such as Dysoxylum 
In Dysoxylum or Hartighsea, in which the styles are 
throughout united and expanded into disc at the summit, the 
centre of which is as usual depressed, the stigmatic surface 
is confined to the lower half of the margin of the disc. 
+ 
¢ 
