PISTILLUM IN CRUCIFER&. 107 
mass interposed between the two sterile carpella, and the 
fact of the endocarp (which is very cellular, hyaline, and 
thick) not entering into the formation, is still more distinct. 
Externally the limits are always distinct, the barren car- 
pella g, being marked out by a line, expanding above into the _ 
styles, the fertile h, by a rim, which becomes more marked 
enc id see Pl. 29.—Fig. 7. e f young, g barren, h 
fertile rim 
Iam aware that an explanation offers itself by merely assum- 
ing the placenta to be the line of union between the 2 car- 
pells, but this will require the lowest part of the sinus of the 
stigmata to correspond to the apex of the style, which is con- 
trary to analogy as well as of venation. The stigmata as they 
are, so obviously correspond to their proper situation, that I 
am willing to look elsewhere for an explanation. I have not 
been able to make sufficiently extended observations with 
very young pistillums. But if my supposition be correct, that 
it is quadri-carpellary, it is a proof that ovuligerous carpella 
may exist without any obvious lamina. It also bears on 
Orchideæ, but is against analogy, since in Cruciferæ and 
Escholtzia, it is the stigma-bearing carpella that bears ex- 
clusively the ovula. And it is probable from the consider- 
ation of Escholtzia, that the imperfection of its stigmata of 
the fertile carpella is connected with the presence of ovula 
or those with perfect stigmata. Neither must I omit ad- 
verting to the probability of the growth of the septum, which 
is an extension from the placenta, being attributable to the 
want of the usual development of the lamina? In Sinapide- 
ous fruits there are obvious traces of four additional com- 
ponent parts, which may be considered as the styles of the 
laminary carpella. See i. Pl. 29.—Fig. 7. [ ? j. k] 
To this structure Escholtzia may be considered as pre- 
senting a transition, although precisely in the opposite direc- 
tion; in that genus there are 4 stigmata, 2 perfect, and 2 
abortive, corresponding to which are 2 dein carpella 
bearing the greater part of the ovula. 
In drawing confirmation of this hypothesis from anomalous 
genera of the natural family, it must be borne in mind, that 
