62 ROYAL SOCIETY OF CANADA 



of Binney,^ as illustrated by a slide, which I owe to his kindness, and also 

 that of the simpler type of JJepidodendra, like Jj. Harcourtii, as described 

 by Williamson, in which no secondaiy or exogenous woody system is 

 developed. My specimen does not, however, show, except in spots, any 

 traces of the inner bark, and the outer or corky layer is. as above stated, 

 pyritized, so that its structure can be seen only imperfectly by reflected 

 light. It is, however, traversed diagonally by the continuations of the 

 leaf-bundles, which show the central bundle of scalariform vessels and 

 two lateral bundles, apparently of hexagonal cells or fibres. 



(3) Fructification. 



The cones or strobiles of I/epidophloios are of the type included in 

 the provisional genus Lepidostrobus ; but are larger and more massive 

 than the cones of ordinary Lepidodendra, and, so far as known, are 

 attached to the sides of the trunk and branches by leafy peduncles or 

 branchlets, either singly or in pairs. In two cases only have I found 

 these cones actually in position. One of these is represented in Plate X. 

 The other I cannot figure, owing, in the first place, to the position of the 

 cone on a short peduncle imbedded in the long leaves of the stem or a 

 large branch, and in the second place to the fact that the original speci- 

 men passed out of my hands many years ago and cannot now be recovered, 

 so that I have only a rough sketch in my note-book of 1851 to represent 

 it. The greater number of the cones which I attribute to my two 

 Acadian species occur separately, and can be recognized only by com- 

 parison of their form and markings. (Plate lY.) None of them show 

 the minute structures, but in one there are rounded bodies which are 

 probably macrospores, scattered in the material occupying the basal 

 portion, so that we may infer Lhat, as in some other Carboniferous trees 

 of Lepidodendroid type, there were maci'ospores below and microspores 

 above. The fertile branchlets, very short in L. Acadianus and much 

 longer in L. Cliftonensis, seem to have been deciduous or easily broken off 

 when mature, leaving tubercles with a central depression, as observed in 

 the fertile branches in the Halonia condition. In the more perfect con- 

 dition of the branches they are concealed by the long and abundant 

 foliage. In branches approaching to the erect position they may be in 

 two ranks or alternate. When by unequal dichotomy there are side 

 branches approaching to a horizontal position, the upper side of the 

 branch may bear alternating cones, while there may be none on the lower 

 side except at the edges, so that this side may appear to bear fruit 

 in two ranks, while on the upper side the arrangement may be irregular 

 or spiral. (Plates V., VII., YIII.) There is no evidence known to me 



1 Publications of Palteontographical Society, 1872. 



