28 



LAMBERSON 



In Stage IV, Pisonia and Cocos dominate the older 

 mixed forest, and Scaevola is nearly shaded out. Morinda, 

 Tournefortia, Guettarda, and Terminalia reach for sun, and 

 Neisosperma may appear. The whole forest is more open, 

 and the undergrowth consists of small Cocos and Pisonia 

 with ground plants of Laportea, Boerhavia, Lepturus, and 

 Portulaca. Sida and Scaeuola are restricted to open areas, 



Suriana and Pemphis occur mostly at the forest margins, 

 and vines become fewer, usually growing only in the sun 

 in clearings and at the outer margins. This stage predom- 

 inated on the main portions of Ikuren and Ananij (Fig. 4). 



In Stage V, Pisonia takes over. Cocos trees become 

 unproductive and gradually die out; other species of trees 

 are confined to the edges of the forest. There is little 



1^ ^'^^ ' 



Fig. 4 Stage IV, older mixed forest. Plants illustrated are predominantly Cocos and Pisonia. 



