46 



CHAVE AND DEVANEY 



'L, lagoon; CH, channel; OS, outer slope. 



tLiving form (followed by color protoplasm); encrusting, test cemented to substrate; attached, holding on to substrate with pseudo- 

 pods; free, not attached to substrate. 



ITotal number of intact specimens in all 42 samples, each containing 1 cc of sand or scraped material. 



The taxonomy to the generic level is based on Loeblich 

 and Tappan (1964, 1974, 1981). Species were identified 

 by Chave using Ellis and Messina (1940 et seq.) and a 

 large number of publications of which Graham and Mili- 

 tante (1959), Barker (1960), Todd (1965), and Belford 

 (1966) were especially useful. 



CILIOPHORA 



Several marine ciliates were collected at Enewetak 

 from reef flat scum (Hirshfield et al., 1957). In addition, 

 Sci;phidium sp. was attached to unspecified invertebrates 

 on the reef, and Ciiclidium sp. was found with habitat 

 unspecified (Hirshfield, op. cit.). Observations made by 

 Patrick Colin from 1979 to 1980 off Runit Islet revealed 

 discolored brownish areas on the lagoon bottom, which 

 were found by Devaney to be formed by high concentra- 

 tions of an undetermined peritrich ciliate that resembles 

 Sc\jphidia. 



During 1966, 42 hymenostomme ciliates were col- 

 lected from the ocean and lagoon sides of nine islets at 

 Enewetak Atoll. Permanent silver-impregnated slides were 

 made of these protozoans from cultures (Thompson and 

 Sellers, 1967), but species were not listed. Subsequently, 

 Thompson (1968) described Cohnilembus verminus from 

 Enewetak. 



Berger (1964) noted that undetermined endocommen- 

 sal ciliates were present in several unspecified regular sea 

 urchins but were absent from at least two species {Euci- 

 daris metuhha and Parasalinia gratiosa) and from irregular 

 urchins and other unspecified echinoderms examined. The 

 classification for ciliophorans follows Corliss (1977). 



OTHER PROTOZOANS 



Fresh and brackish water protozoans at Enewetak 

 revealed mainly amoeboid types from nearly 100 water 

 and soil sites on nine islets. Additional fish, gecko, rat and 

 bird blood, and other tissues were examined for protozoan 

 parasites (Dillon and Kasweck, 1970). 



NEW ENEWETAK RECORDS 



In 1981, 42 samples were studied at Enewetak 

 immediately after being collected from the lagoon at 



depths of from 1 to 61 m, in the channels from 1 to 

 33 m, and on the outer slopes from 1 to 400 m. Of the 

 new Enewetak records, 23 species were represented by 

 one to three specimens and 14 species were fairly com- 

 mon but were obtained by methods not used during the 

 1946 collections (hard surfaces were scraped by hand to 

 depths of 33 m, and rocks were brought up from deep 

 areas by the submersible Makili'i). The final 10 species 

 were common in sediment samples and were not reported 

 by Cushman et al. (1954). 



Twenty-nine species listed by Cushman et al. (1954) 

 were not found by the senior author. Twenty-seven of 

 these species were listed as rare or from the 914 m sta- 

 tion (op. cit). Two species, Hauerina serrata and 

 Asfrononion tumidum, were listed as common in lagoon 

 sediments (op. cit.). 



ACKNOWLEDGMENTS 



We acknowledge and are grateful to H. I. Hirshfield, 

 New York University, for permission to include the records 

 of ciliates given in his 1957 report to MPRL. J. Resig at 

 the University of Hawaii made valuable suggestions and 

 corrections in the Foraminifera part of this paper. The 

 facilities at Enewetak and material obtained by the Hawaii 

 Undersea Research Laboratory's submersible Makali'i 

 were made available to both authors through grants to 

 Patrick Colin and John Harrison of MPRL from NOAA's 

 National Undersea Research Program, the Department of 

 Energy, and the Defense Nuclear Agency. 



REFERENCES 



Barker, R. W., 1960, Taxonomic Notes on the Species Figured 

 by H. B. Brady in his Report on the Foraminifera Dredged by 

 the HMS Challenger During the Years 1873-1876, S.E.P.M. 

 Spec. Pub. 9. Tulsa, Oklahoma, 238 pp. 



Belford, D. J., 1966, Miocene and Pliocene Smaller Forami- 

 nifera from Papua and New Guinea, Austr. Bur. Min. Res. 

 Geo/. Geoph. Bull. 79. 306 pp. 



Berger, J., 1964, Mid Pacific Research Laboratory Annual Rept., 

 FY 1 963-64, 3 pp. 



Cole, W. S., 1957, Larger Foraminifera from Eniwetok Atoll Drill 

 Holes, U. S. Geol. Surv. Prof. Pap. 260 V. pp. 743-784. 



