154 



SAMUELSON AND NISHIDA 



TABLE 2 

 Collectors of Insects and Related Groups on Enewetak Atoll 



Name 



Month/Year 



Islets 



Allen, R. 

 Bohart, R. 

 Bryan, E. H. 



Jr. 



Bushman, J. B. 



Cheng, L. 

 Dybas, H. S. 

 Edgar, S. A. 

 Fosberg, F. R. 



Hagen, E. 

 Jackson, W. B. 

 Knudsen, J W. 



Lamberson, P. 

 Morrison, J. 



Oakley, R. G. 



Oshiro, Y. 

 Parker, D. 

 Reish, D. J. 

 Townes, H. K. 



Tuthill, L. D. 



Wheeler, M. R. 

 Young, F. N. 



Aug. 1945 

 Jan. 1945 

 Aug. 1944 

 Feb. 1975 



Mar. -May 1962 



Sept. 1975 

 Nov. -Oct. 1944 

 Nov. -Oct. 1944 

 May 1946 

 May 1951 

 Nov. 1944 

 1964-67 (summers) 

 Apr. 1965 

 Aug. 1967 



Apr-June 1946 



May 1946 



Dec. 1950-Jan. 1951 

 Mar. -Apr. 1962 

 July 1956 

 May 1946 



Aug. Sept. 1956 

 Jan. -Feb. 1957 

 July 1957 

 Sept. 1957 

 Aug. 1958 

 Aug. -Sept. 1955 

 May 1945 



Enewetak, Runit 



Enewetak? 



Enewetak, Enjebi, Japtan, Medren 



Alembel, Ananij, Bijile, Enewetak, 



Enjebi, Ikuren, Japtan, Lojwa, 



Medren, Runit 

 Aej, Enewetak, Enjebi, inedral, 



Japtan, Jinimi, Medren, Mijikadrek, 



Ribewon 

 Enewetak, Ikuren, Japtan, Medren 

 Enewetak, Japtan 

 Enewetak, Japtan 

 Japtan 

 Enjebi 



Enewetak, Japtan 

 Enjebi, Runit 

 Enewetak 



Enjebi 



Biken, Ikuren, Kidrinen, Louj, Lujor, 



Mut 

 Aomon, Enjebi, Japtan (and others 



with Townes?) 

 Bokombako, Elugelab, Enjebi 

 Aej 



Enewetak 

 Aomon, Enewetak, Enjebi, Ikuren, 



Japtan 

 Ananij, Ikuren, Japtan, Medren 

 Japtan, Medren 

 "Nan" 

 Medren 

 Japtan 



Enjebi, Ikuren, Japtan, Medren 

 Medren 



Enewetak during the same periods. Hence, the indigenous 

 {including endemic) biota of Enewetak — the species that 

 represent the true unadulterated character of the 

 atoll — probably suffered profoundly through both physical 

 impacts on the environment and biological impacts, 

 including competition for space and resources from adven- 

 tive forms. 



In view of the disturbances to Enewetak Atoll, it seems 

 relevant to note the times and places where species were 

 collected or observed, thereby providing a summary of the 

 first four decades of entomological records for the atoll. 

 This information is provided in the section on Collection 

 Records. 



Fauna 



Townes (1946) estimated that a large dry atoll like 

 Enewetak might have 500 species of insects. This estimate 



could be somewhat high. The indigenous insect fauna of 

 Enewetak would be expected to be less than that of Arno 

 Atoll. The fauna of Arno was also estimated at around 

 500 species (Usinger and LaRivers, 1953), and at least 

 296 species have already been recorded for that atoll 

 (Gressitt, 1954). Cole (1949, 1951) listed 56 species of 

 insects collected from Bikini Atoll in 1947. Bikini lies in 

 the same zone as Enewetak and would be expected to sup)- 

 port a nearly identical fauna. Cole's survey was the only 

 one conducted for Bikini and, unfortunately, it occurred 

 after applications of DDT on the islets and after the first 

 atomic bomb tests there. These perturbations must have 

 had severe effects on the fauna and on Cole's results. 

 Cole's lists were admittedly incomplete; however, 35 

 species listed by him, including 12 species of ants, have 

 not yet been reported for Enewetak. Most of these 35 

 species probably also occur or have occurred on 

 Enewetak. 



