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SAMUELSON AND NISHIDA 



fauna comprises 89 species, with endemism about 81%. 

 Four species are restricted to atolls; all are in 

 Campiilomma: the two mentioned above, plus C. utohiensis 

 Carvalho (Ulithi) and C. wakeana Schuh (Wake). 



Nabidae (Gross, 1963): Marshalls 1, Enewetak 

 1 — Nabis capsiformis Germar is a cosmopolitan species. 

 Micronesian fauna comprises four species of which one is 

 endemic to the Carolines (Palau, Ponape). 



Pentatomidae (Ruckes, 1963): Marshalls 2, Enewetak 

 2 — The Plati/nopus we list for Enewetak is restricted to 

 Melanesia and Micronesia, and the Oechalia we list is of 

 Australian-Pacific distribution. Micronesian fauna com- 

 prises 33 species, with endemism about 55%. All of these 

 species include high islands in their distributions. 



Reduviidae (Wygodzinsky and Usinger, 1960): 

 Marshalls 1, Enewetak — The only species recorded from 

 isolated atolls is the one reaching the Marshalls, Phiaria 

 insolida (B. White), which is Philippines-Pacific in 

 distribution. Micronesian fauna comprises 30 species, with 

 endemism about 67%. 



Veliidae: Marshalls 1, Enewetak — Halouelia mahtima 

 Bergroth is marine and mainly of Australian-Indonesian 

 distribution; it is also reported for Bikini (Cole, 1951). 



Homoptcra 



This is the only large insect order reasonably stutied in 

 its entirety for Micronesia. Some of the families are well 

 represented on Micronesian islands and have a high pro- 

 portion of endemic taxa. The following families were 

 treated for Micronesia but representatives are not reported 

 for the Marshalls: Alcyrodidae (Takahashi, 1956), Cercopi- 

 dae (Synave, 1957), Cicadidae (Esaki and Miyamoto, 

 1975), and Membracidae (Kato, 1960). 



Aphididae (Essig, 1956): Marshalls 4, Enewetak 2 — All 

 are widespread or cosmopolitan species also reported for 

 Hawaii. An additional aphid reported for the Marshalls is 

 Hiisteroneura setariae (Thomas) (Sugerman, 1972a). 

 Micronesian fauna comprises 16 species, mostly 

 widespread; endemics include one species and one sub- 

 species of Malayan or Oriental affinities. 



Cicadellidae (Linnavuori, 1960, 1975): Marshalls 10, 

 Enewetak 7 — The species we list for Enewetak are 

 widespread in both hemispheres (1), of New World-Pacific 

 distribution (1), Oriental-Pacific (3), Australia-Pacific (1), 

 or limited to Oceania (1). At least one is reported for 

 Hawaii: Recilia affinis (Osborn). Baldutha lucida (Butler) is 

 reported for Arno and Ebon but not Enewetak; it is of 

 New World-Pacific distribution. Two undetermined cicadel- 

 lids are reported for Kwajalein by Sugerman (1972a, 

 1979): Empoasca, a genus well represented in the Pacific 

 and Carneocephah, a New World genus not included in 

 Linnavuori's treatment. Micronesian fauna comprises 87 

 species, with endemism about 66%. 



Cixiidae (Fennah, 1956, 1971): Marshalls 2, Enewetak 

 — Both Myndiis apicalis (Metcalf) and M. praecanus Fen- 

 nah are endemic to the central eastern Carolines and 

 Marshalls. Micronesian fauna comprises 29 species, all 

 endemic. 



Coccoidea (Beardsley, 1966, 1975): Marshalls 36, 

 Enewetak 7 — These are mostly species of widespread dis- 

 tribution of which 25 also occur in Hawaii. Seven species 

 are restricted to Micronesia; of these, five are widespread 

 in Micronesia and two arc endemic to the eastern Caro- 

 lines and Marshalls. Many of these scale insects are com- 

 mon on cultivated plants and have become widely dis- 

 tributed. All of the species reported for Enewetak also 

 occur on Kwajalein, and five of the seven also occur in 

 Hawaii. Micronesian fauna comprises 150 species, with 

 endemism about 23%. 



Delphacidae (Fennah, 1956, 1971): Marshalls 9, 

 Enewetak 1 — These species are restricted to Micronesia 

 (2), Micronesia plus Hawaii (1), or mainly to Oceania plus 

 continents and continental islands bordering the western 

 Pacific (6). The Sogatella we list for Enewetak is reported 

 for Australia, Philippines, and Micronesia. Of the restricted 

 taxa, Ug\/ops kinbergi manias Fennah is endemic to the 

 Marshalls and occurs on both island chains and 

 U. superciliatus Fennah is limited to the eastern Carolines 

 and Marshalls. Micronesian fauna comprises 63 species 

 with endemism about 44%. Three species are endemic to 

 atolls elsewhere: the Gilberts, Wake, and Ocean Island, a 

 raised atoll. 



Derbidae (Fennah, 1956, 1971): Marshalls 3, Enewetak 

 1 — Species or subspecies restricted to Micronesia of 

 which the Lamenia we list for Enewetak is endemic to the 

 Marshalls; Flaccia dione Fennah is limited to Caroline 

 atolls, Kosrac, and Marshalls; and Smezeyia zephrus Fen- 

 nah occurs through much of Micronesia. Lamenia and 

 Siuezeyia possibly have Philippine affinities, while Flaccia 

 has ties to Melanesia (Fiji). Micronesian fauna comprises 56 

 species, with endemism about 95%. Derbids are well 

 represented on Caroline atolls, with 20 species of which 

 four are endemic to atolls; another species is endemic to 

 the Gilberts. 



Psyllidae (Tuthill, 1964): Marshalls 1, Enewetak 

 — Mesohomotoma hibisci (Froggatt) ranges from Australia 

 and continental islands bordering the western Pacific to 

 Oceania where it is widespread; it is reported for 

 Kwajalein (Sugerman, 1972a). Micronesian fauna 

 comprises 22 species, with endemism about 68%. Two 

 additional psyllids besides M. hibisci occur on Caroline 

 atolls but are not reported for the Marshalls. 



Coleoptera 



Most of the large families of beetles have been sur- 

 veyed. The outstanding exception is the Curculionidae or 

 weevils, a family well represented on Pacific islands, with 

 only the Guam components of the Micronesian fauna ade- 

 quately studied. A few species are reported for the 

 Marshalls but not Enewetak. Families monographed for 

 Micronesia, but not yet reported for the Marshalls, include 

 the Cantharidae (Wittmer, 1958), Cryptophagidae (Michi- 

 taka Chujo, 1970), Cybocephalidae (Endrody-Younga, 

 1971), Endomychidae (Strohecker, 1958), Lampyridae 

 (Wittmer, 1958), Malachiidae (Wittmer, 1958, 1970), Pla- 



