INSECTS AND ALLIES (ARTHROPODA) 



163 



typodidae (Wood, 1960), Prionoceridae (Wittmer, 1958), 

 Rhysodidae (Bell and Bell, 1981), and Scaphidiidae (Lobl, 

 1981). 



Anobiidae (Ford, 1958): Marshalls 1, Enewetak 

 — Lasioderma serricorne (Fabricius), the cigarette beetle, 

 is reported for Kwajalein; it is a cosmopolitan species. 

 Microncsian fauna comprises nine species, with endemism 

 about 67%. 



Anthicidae (Werner, 1965); Marshalls 3, Enewetak 

 1 — All are species of Anthicits and are confined to the 

 Pacific Basin. Anthicus vexator Werner, reported for 

 Enewetak, occurs on Caroline atolls and the Marshalls and 

 Gilberts. It is also reported for the Phoenix Islands and 

 Hawaii. Anthicus insuhhs Werner is restricted to 

 Micronesia and A. oceanicus LaFerte is widely distributed 

 in Micronesia, extending to Polynesia, Ryukyus, and possi- 

 bly into the Indian Ocean. Micronesian fauna comprises 

 nine species, with endemism about 44%. 



Anthribidae: Marshalls 1, Enewetak — Araecerus sp. 

 is reported for Kwajalein (Sugerman, 1972a) and is prob- 

 ably one of several species widespread in the Pacific. The 

 genus is mainly Oriental-Pacific in distribution. 



Bostrychidae (Michio Chujo, 1958): Marshalls 1, 

 Enewetak — Dinoderus bifoueolatus (Wollaston), widely 

 distributed in warmer regions, is reported for Kwajalein. 

 Micronesian fauna comprises 10 species, with none 

 endemic; most are widespread. 



Carabidae and Cicindelidae (Darlington, 1970); 

 Marshalls 5, Enewetak 1 — These species are of 

 Oriental-Pacific distribution (1) or Indo- Australian-Pacific 

 (1), of Neotropical affinities (2), or possibly restricted to 

 Oceania (1). The Egadroma we list for Enewetak belongs 

 to the second category. Two species were not treated by 

 Darlington; they are Callida insuhris Boheman and 

 Selenophorus sp.; both were reported for Jaluit by Schnee 

 (1904). Micronesian fauna comprises 49 species, with 

 endemism about 27%, representing 11 different stocks, 

 most resulting in only a single endemic species. Curiously, 

 one cicindelid, Therates labiatus (Fabricius), ranging from 

 Melanesia to the Philippines, also occurs on Nukoro, one 

 of the Caroline atolls, but nowhere else in Micronesia. 



Cerambycidae (Gressitt, 1956): Marshalls 12, Enewetak 

 1 — These species are mostly restricted to Micronesia and 

 have Oceanian-Malayan affinities; close relatives may 

 extend to the Indian Ocean, but they are generally absent 

 on continental Asia. Of these, five are endemic to the 

 Marshalls: two Ceresium are restricted to single atolls, C. 

 unicolor marshallum Gressitt (Majuro) and C. robustum 

 Gressitt (Arno); Prosoplus major Gressitt and P. hibisci 

 Gressitt are from various atolls, also in the eastern chain 

 (Ratak); and Oopsis marshallensis Gressitt is the most 

 widespread of the endemics, occurring on both island 

 chains. The Prosoplus we list for Enewetak occurs in the 

 eastern Carolines (Ponape) and Marshalls. Only Si>bra 

 alternans (Wiedemann) ranges beyond Micronesia; it is also 

 reported for Indonesia, Philippines, and Hawaii. Microne- 

 sian fauna comprises 105 species, with about 87% re- 

 stricted to Micronesia, including 7% endemic to atolls or 



low islands. Correspondence of species occurring on the 

 different Micronesian atoll groups Is fairly low, with three 

 of 12 species from Caroline atolls also present in the 

 Marshalls and one of the four species from the Gilberts 

 likewise found in the Marshalls. Most of the atoll species in 

 the Carolines also occur on adjacent high islands; however, 

 Sciadella atolhrum Gressitt is restricted to atolls. One of 

 the four Gilberts species, S^ibra catahna Gressitt, is 

 endemic there. 



Chrysomelidae (Gressitt, 1955): Marshalls 3, Enewetak 

 1 — These species are either confined to Oceania and their 

 affinities are Melanesian-Malayan (2) or they are 

 Oriental-Pacific in distribution (1). Aphthona bicolorata 

 Jacoby is associated with euphorbiaceous plants and is 

 fairly common in parts of Oceania, including Enewetak. 

 Brontispa chali^beipenms (Zacher) is associated with the 

 coconut palm and had a rather restricted range, within the 

 eastern Carolines (Fonape, Kosrae) and the Marshalls 

 before it reached Hawaii. Pagria signata (Motschulsky) 

 belongs to the second category. Micronesian fauna 

 comprises about 38 species, with endemism about 66%. 

 None of the species is restricted to atolls. 



Cleridae: Marshalls 1, Enewetak — One cosmopolitan 

 species, Necrobia rufipes DeGeer, the red-legged ham bee- 

 tle, is reported for Jaluit (Schnee, 1904). 



Coccinellidae (Chapin, 1965): Marshalls 8, Enewetak 

 4 — These species either are found on continents, Asia 

 and/or Australia (4), or are confined to the Pacific Basin, 

 with two restricted to Micronesia and two also found on 

 subcontinental islands of the western Pacific. One of the 

 restricted species, Sci/mnus uncinus Chapin, is limited to 

 the Marshalls and Gilberts. One additional species, from 

 India, was released in the Marshalls (Majuro, Uliga) and 

 Guam but was not recovered. Micronesian fauna comprises 

 48 species, with endemism about 46%. 



Cucujidae: Marshalls 1, Enewetak — The cosmop>oli- 

 tan Ort^zaephilus surinamensis (Linnaeus), the saw-toothed 

 grain beetle, is reported for Kwajalein (Sugerman, 1972a). 



Curculionidae (Zimmerman, 1942, 1948, 1964): 

 Marshalls 6, Enewetak — These species are possibly 

 cosmopolitan {Sitophilus sp.), of Indonesian-Pacific affini- 

 ties (Oxt;dema sp.), restricted to Oceania (3), or undeter- 

 mined (1). Of the restricted species, Sphenophorus sulcipes 

 Karsch is described from the Marshalls; Plafysimus insuhris 

 (Boheman) is reported for Polynesia, Fiji, the Marshalls and 

 Gilberts; and Trigonops hirsute Zimmerman, described 

 from Guam, is reported for Kwajalein. Weevils of Guam 

 and the Marianas have received attention by Zimmerman 

 (1942, 1948). 



Dermestidae (Beal, 1961): Marshalls 3, Enewetak 

 1 — All of the species reported for the Marshalls are also 

 found in Hawaii, and none is restricted to the Pacific. The 

 Dermestes we list for Enewetak is cosmopolitan and 

 widespread in Micronesia. Orphinus terminate (Sharp) is of 

 Malayan-Pacific distribution and Trogoderma anthrenoides 

 (Sharp) is a Neotropical species that has been established 

 in Hawaii and has also reached Saipan and Kwajalein. 

 Micronesian fauna comprises 12 species, mostly wide- 



