164 



SAMUELSON AND NISHIDA 



spread; only one species, Orphinus nesioticus Beal, from 

 Guam, is endemic. 



Elateridae (Van Zwaluwenburg, 1948, 1957; Ohira, 

 1971): Marshalls 11, Enewetak 2 — All identified species 

 are of Oceanian distribution. The two species we list for 

 Enewetak are either reported for New Britain, eastern 

 Carolines, and Marshalls (Simodacti^lus) or range through 

 Micronesia and Polynesia (Conoderus). Two other species 

 are endemic to the Marshalls and are restricted to single 

 atolls: Melanoxanthus lariversi Van Zwaluwenburg from 

 Arno and Sirnodacfylus marshallensis Ohira from 

 Kwajalein. Two unidentified elaterids, Ampedus sp. and 

 Anchastus sp., are reported for Kwajalein (Sugerman, 

 1972a, 1979). Microncsian fauna comprises 70 species, 

 with endemism about 71%. 



Histeridae: Marshalls 1, Enewetak — Plati^soma sp. is 

 reported for Jaluit (Schnee, 1904). The genus is well 

 represented in the Indo-Malayan area, but some species 

 occur in the New World. 



Hydrophilidae: Marshalls 1, Enewetak — At least one 

 species, Dacfylosternum abdominale (Fabricius), is reported 

 for the Marshalls (Sugerman, 1972a); it is possibly the 

 same as one or both of the C\jclonotiim spp. reported by 

 Schnee (1904) for Jaluit. The species is widely distributed 

 in the New World and is established in Hawaii. 



Mycetophagidae (Michitaka Chujo, 1970): Marshalls 2, 

 Enewetak 1 — The Tijphaea we list for Enewetak is a 

 cosmopolitan species. Litargus uestitus Sharp, reported for 

 Arno and Jaluit, is an Oceanian species found throughout 

 Micronesia and parts of Polynesia. Both species are 

 reported for Hawaii. Microncsian fauna comprises three 

 species, with none endemic. 



Nitidulidae (Gillogly, 1962): Marshalls 14, Enewetak 

 6 — These are mostly widespread species, but one is re- 

 stricted to Micronesia and Polynesia; six are cosmopolitan; 

 three are found on various continents; one is also from 

 Australia; two are also from New Guinea; and one occurs 

 in Polynesia, Philippines, and Sri Lanka. Micronesian fauna 

 comprises 39 species, with endemism about 33%. None is 

 restricted to atolls; correspondence of species among dif- 

 ferent atoll groups is fairly high with six of eight of the 

 Caroline atolls species and seven of the nine Gilberts spe- 

 cies also found in the Marshalls. 



Oedemeridae (Macnamara and Arnctt, in preparation): 

 Marshalls 8, Enewetak 6 — These species are more or less 

 of Malayan-Oriental distribution. Of these, only one is 

 rather restricted, occurring only in the Marshalls, Gilberts, 

 and Ocean Island. The remaining seven tend to be 

 widespread in Micronesia and include some that also occur 

 on Melanesian and Polynesian islands. Micronesian fauna 

 comprises 19 species, with most restricted to Micronesia. 



Propalticidae (John, 1960, 1971): Marshalls 1, 

 Enewetak 1 — Propalticus insularis John is restricted to 

 Micronesia. Micronesian fauna comprises eight species, 

 with endemism 63%. 



Scarabaeidae (Cartwright and Gordon, 1971): Marshalls 

 6, Enewetak — These species are of the subfamily Apho- 

 diinae and arc either restricted to Micronesia (2); nearly 



restricted, occurring also in Samoa (1); found in the 

 Malayan and/or Melanesian subregions (2); or have a gen- 

 eral Old World distribution (1). Micronesian fauna 

 comprises 27 species (10 are aphodiines), with endemism 

 about 37%. Of the endemic species, four are aphodiines, 

 four are melolonthines, one is a ruteline, and one is a 

 dynastine. None is restricted to atolls. Both species occur- 

 ring on Caroline atolls (aphodiines) also occur in the 

 Marshalls. One of the two Gilberts species, also an apho- 

 diine, occurs in the Marshalls. The other species, a dynas- 

 tine, Papuana huebneri (Fairmaire), is the only Micronesian 

 record for that Melanesian group. 



Scolytidae (Wood, 1960): Marshalls 9, Enewetak 

 2 — Of these species, only one is endemic to a more or 

 less restricted area, Kosrae and the Marshalls including 

 Enewetak; one species reaches Hawaii; one is Malayan; 

 two are cosmopolitan; and the rest tend to be widespread 

 in Micronesia and also occur in warmer regions bordering 

 the western Pacific. Microncsian fauna comprises 53 

 species, with endemism about 28%. 



Staphylinidae: Marshalls 2, Enewetak — Two rove 

 beetles arc reported for Kwajalein by Sugerman (1972a): 

 Lispinus impressicollis Motschulsky, ranging throughout the 

 Old World tropics and extending into the Indian and 

 Pacific Oceans, and a species questionably placed in 

 Medon, a widespread genus with many species. 



Tencbrionidae (Kulzer, 1957): Marshalls 16, Enewetak 

 2 — Of these species, two are endemic to the Marshalls; 

 one also occurs on Kosrae, eastern Carolines; three are 

 more widespread in Micronesia; three are Oceanian; two 

 arc Malayan-Pacific and comprise the Gebieniella and 

 Gonocephalum that we list for Enewetak; one is Oriental; 

 and four are cosmopolitan. Of the species endemic to the 

 Marshalls, Tagalus angustus Kulzer is restricted to Arno 

 and Chariotheca costata Kulzer is restricted to Arno and 

 Ailinglapalap. Micronesian fauna comprises 92 species, 

 with about 62% restricted to Micronesia, including 4% 

 endemic to atolls and low islands. Correspondence of 

 species between atoll groups is fairly low, with four of 13 

 Caroline atolls species and one of the three Gilberts 

 species also occurring in the Marshalls. One of the Caro- 

 line atolls species, Bradxjmerus faraulepensis Kulzer, is 

 endemic to a single atoll (Faraulep). 



Strepsiptera 



Micronesian fauna unstudied. 



Neuroptera 



Only the Chrysopidae has been reported for the 

 Marshalls. Two other families treated for Micronesia are 

 still unreported for the Marshalls: Hemerobiidae (Car- 

 penter, 1961) and Myrmeleontidae (Adams, 1959). 



Chrysopidae (Adams, 1959): Marshalls 2, Enewetak 

 1 — Both species are widespread in Micronesia and are 

 also of Australian-Malayan distribution. Chri>sopa ramburi 

 Schneider, reported for Enewetak, is the senior synonym 



