Law of Proenvironivient 223 



vironal responses. But in all nucleate or chroniatin-bearing 

 plants, and in the simpler non-ganglionic animals that are also 

 chromatin-bearing, the perceptive or cognitic energy that we 

 beheve to be associated with and to traverse the chromatin 

 substance acts intrinsically so as to link up or combine into 

 one resultant response the separate responses of two or more 

 energizing stimuli, and so to proenviron a course that is the re- 

 sultant of ally and that is satisfying to the organism. 



But, '^'ith the gradual formation and appearance in nerve 

 cells of ganglionic substance or neuratin, the still higher cogitic 

 energ^^ that we consider to be associated wath and to traverse 

 its substance acts intrinsically so as to absorb two or more 

 sets of distinct resultant responses, and then link up or com- 

 bine these into one compound resultant response that is satis- 

 fying to the ganglionated organism. Hence result, as already 

 stated, the responses that we associate with memory, mind, 

 intelligence, intellectuality, and morality, that are due to 

 combined complex action and reaction of bio tic, cognitic, 

 and cogitic energies acting in an increasingly more and more 

 complex manner through the protoplasm, chromatin, and 

 neuratin (if for convenience we may adopt such a term) so as 

 to exhibit (a) simple protoplasmic responses, or (b) resultant 

 chromatin responses, or (c) compounded resultant ganglionic 

 or reflex responses. 



An attempt may now be made, therefore, to trace the pos- 

 sible gradual segregation and evolution of nerve cell or gan- 

 glionic substance, beginning with the more specialized Protozoa 

 and the simpler Metazoa, and then passing upward through 

 the higher groups of the animal kingdom. Two of the higher 

 groups of Protozoa seem to furnish a key to initial conditions. 



Amongst the Hcemoflagellata, according to the researches of 

 Schaudinn, Leger, and Prowazek, each young cell-individual 

 shows a single nucleus. But during cell maturation the nucleus 

 divides, and one of these redivides, so as to give rise to three 

 nuclei. One of the three has been called the trophonucleus, 

 as it seems to preside over or govern the general sensory and 

 trophic functions of the cell. But without desiring to coin 

 new words unnecessarily, the writer will call it the cognito- 



