108 THE ROYAL SOCIETY OF CANADA 



single vascular strand and branched dichotomously (and so differed 

 from those of Stigmaria proper, fide Potonié). "Stigmaria" is rare 

 in Devonian rocks, but one of the commonest fossils of the Coal 

 Measures. 



Strobile, Plate I, Fig. 3; Plate III, Fig. 2. 



"There is considerable difference in the strobiles or cones of the 

 several species of fossil plants referred to Bothrodendron. It ha;- 

 been generally agreed that the cone from Kiltorcan described as 

 Lepidostrohus Bailyana is the fruiting strobile of Cyclostigma Kiltor- 

 kense. It is terminal to the stem, and obconical in shape. From one 

 specimen it appears that the axis was hollow, but contained tissue 

 that was removed in the process of fossilization. In the space en- 

 closed by the cylindrical axis is a thin irregularly shaped carbonized 

 body, whose nature is not obvious; a longitudinal section is shown in 

 the breaking of the stone, revealing a carbonized partition, which 

 stretches across the central space of the cone. Numerous impressions 

 in the slab show the sporophylls from many points of view. The 

 sporophyll is 20 cm., or more in length, has a thick spatulate, or 

 sub-obtriangular fertile base, 16 m.m. long and 1 to 1-5 m.m. broad 

 at the point of attachment, 3 to 5 m.m. at its upper end; and a long 

 narrow awlshaped appendage or upper part, 2 m.m., wide at its point 

 of connection with the distal end of the fertile base. These two parts 

 do not lie in one plane, but are generally at an angle, so that the free 

 end of the sporophyll in the cone was directed outward as well as 

 upward. The fertile base shows a median groove on the under or 

 abaxial side, and a pronounced ridge on its upper side, probably fitting 

 into a groove on the underside of the sporophyll, immediately above 

 it in the cone. On either side of the ridge there is a line or slight 

 groove (parichnos continuation ?) Deprived of its fertile proximal 

 end, the sporophyll appears similar to an ordinary leaf. 



"It is on the upper or abaxial side of the megasporophyll that the 

 megaspores are borne. They are 1 mm. in diameter, and rounded, 

 or as sometimes seen, pyriform bodies. Schimper's figure of the 

 sporophyll of L. bayliana shows numerous megaspores, each with a 

 triradiate mark, indicative of fan origin by division of the megaspore 

 mothercell into four spores. As the drawing was made ad natiiram, 

 and it is stated that though one may safely assume that the tetrahid 

 division took place, I have found nothing comparable to Schimper's 

 figure in the very many sporophylls I have examined. 



"Occasionally the spores have appeared arranged as if with a 

 common origin. They are found more or less in two rows on each 

 side of the midrib, from 20 to 30 in number in each sporophyll. De- 



