BORAGINACEAE 393 
120. BORAGINACEAE! (BoRAGE or ANONANG FAMILY) 
Herbs, shrubs, or trees. Leaves alternate, mostly entire, simple, stipules 
none. Inflorescence mostly terminal, of forked, scorpoid cymes, sometimes 
spike-like, or corymbose. Flowers perfect. Cabhyx inferior, subentire, or 
5-, rarely 4- to 8-lobed or toothed, usually persistent. Corolla tubular, 
4- to 8-, mostly 5-lobed. Stamens as many as the corolla-lobes, alternate 
with the lobes, inserted on the tube. Ovary superior, 2-celled, 2-ovuled, 
or 4-celled, 1-ovuled; style terminal or from between the ovary-lobes. 
Fruit a small drupe or divided into 2 or 4 nutlets. 
Genera 85, species more than 1,200 in most parts of the world, 10 genera 
and 21 species in the Philippines. 
1. Shrubs, trees, or climbing vines. 
2. Erect shrubs or trees. 
3. Styles single, twice forked............... EP RLS Ae 1. Cordia 
Re ee ee Ns So”: en. % 2. Ehretia 
Ri Reman, LES Ree RI Set I Oe 3. Tournefortia 
1. Herbs. 
Di Rpeaicmm GG “Heariated - 220022252 Ae ee 28 ke. 4. Heliotropium 
2. Inflorescence not scorpoid. 
8. Erect, branched, the flowers of medium size in open leafy racemes 
Sp BS Be, Oe Shy RCL Dee aoe eae 5. Trichodesma 
3. Prostrate, spreading, the flowers small, axillary, solitary or in very 
NGG VSILOE: BEB OETE OOS BO ne OF os IE Raed 6. Coldenia 
1. CORDIA Linnaeus 
Trees or shrubs. Leaves alternate, petioled, often repand or undulate. 
Flowers in terminal dichotomous corymbs, ebracteate, polygamous, those 
with perfect flowers smaller than those with male ones. Calyx tubular or 
bell-shaped, teeth short, often irregular, in fruit accrescent and persistent. 
Corolla funnel-shaped, 4- to 8-lobed, the lobes spreading, white or yellow. 
Stamens as many as the corolla-lobes; filaments often hairy at the base. 
Ovary 4-celled, glabrous; style bipartite, the arms again cleft. Drupe ovoid 
or ellipsoid, stone bony, 1- to 4-celled, usually only 1 cell with a seed. (In 
honor of E. Cordus, an early German botanist.) 
Species about 180, in all tropical countries, 3 in the Philippines. 
1. ©. myxa L. (C. blancoi Vid.)). Anonang (Tag.) 
A tree 5 to 10 m high, glabrous or nearly so. Leaves ovate to oblong- 
ovate or elliptic-ovate, 6 to 15 cm long, entire or somewhat undulate-repand, 
shortly blunt-acuminate, base acute, or subcordate-rounded. Inflorescence 
corymbose, axillary and terminating short lateral branches, lax, 5 to 10 
em long. Flowers white or yellowish-white, sessile, about 7 mm _ long. 
Calyx ovoid. Corolla-tube not longer than the calyx, the lobes spreading, 
throat and stamens villous, the lobes reflexed. Drupe ovoid, yellowish- 
white, 10 to 13 mm long, pulp rather scanty, stone hard, the persistent 
calyx broadly funnel-shaped, enlarged, about 8 mm in diameter. (Fl. Filip. 
pl. 48, C. sebestena.) 
In thickets, Pasay, etc., fl. Apr.-June; widely distributed in the Philip- 
pines. India to Malaya. 


1For a consideration of all the known Philippine species of this family 
see Robinson, C. B., “Philippine Boraginaceae.” Philip. Journ. Sci. 4, 
(1909) Bot. 687-698. 
