FEEDING 



287 



hence stretched. The hning contains fundus glands, and both pepsin and 

 hydrochloric acid have been found in this compartment. Japanese 

 scientists, in particular, have studied the quantity and the effects of these 

 pepsins with a view to applying them to therapeutic purposes. In Europe, 

 pepsin forms the basis of various pharmaceutical preparations, and is 

 generally derived from the stomachs of cattle, but Japan, lacking a 

 pastoral economy, is forced to rely partly on whales for her pepsin 

 supplies. 



Apart from pepsin, the second compartment also contains lipase, a 

 fat-digesting enzyme which, though generally secreted by the pancreas, 

 may also be formed in the stomachs of terrestrial animals and of Carnivores, 

 in particular. H. J. Ketellapper, who examined Blue and Fin Whale 

 stomachs inter alia for the presence of enzymes, shows that only small 

 traces of lipase were present in them. 



The lining of the third compartment, the pyloric stomach, is generally 

 reddish-brown in colour, and looks as velvety as the second. While it may 

 have a few fine folds, it is often very smooth, and contains a large number 

 of normal pyloric glands. 



If we compare the Cetacean stomach with that of other mammals. 



Figure 16 j. Whenever the stomacli oj a whale is accidentally cut open, the krill spills out over 

 the deck. {Photograph: H. W. Sytnons, London.) 



