556 LXXXIX. PEDALTNEiE. 



superposed in a single row or rarely solitary. Style filifonnj witli as^many 

 stigmatic lobes as carpels. ' Fruit diy, hard and indehiscent or opening in 

 valves. Seeds with a' thin testa. Albumen scanty or none. Embryo 

 straight, with a very short radicle. — Herbs. Leaves all or at least the lower 

 ones opposite. Flowers solitary in the axils of the floral leaves or bracts, the 

 upper ones often forming a terminal raceme, with or without bracteoles. 



A small Order, dispersed over the tropical and subtropical regions both of the New aud 

 the Old World, the only Australian genus extending to the Eastern Archipelago. De Can- 

 dolle's arraiigementj including Sesameje and Pedaleae in one Order, appears to be far the 

 most natural ; thus forming a small group connected on the one hand with Gcsueriaceffi and 

 Bignoniaceie by their flowers, and on the other hand with Verbenaceee by their ovary or 

 fruit divided into twice as many cells as carpels, 



1. JOSEPHINIA, Vent. 



Calyx divided to the base into 5 segments. Corolla tubular, the 

 lobes spreadint^, short, the lowest rather larger than the others. Stamens 

 .didynanious, included in the tube; anther-cells parallel, the conncctivum 

 usually tipped with a small gland. Ovary of 4, 6 or 8 cells, each with 1 

 ■erect ovule; stigmatic lobes 2, 3 or 4. Fruit hard and indehiscent, armed 

 with conical prickles, shortly or not at all beaked. Seeds 1 in each cell, 

 oblong, erect. — Herbs with the habit of Sesamiim, Leaves opposite, entu'C, 

 toothed or divided. Mowers in the upper axils on short pedicels without 

 bracteoles. 



The genus extends into the Archipelago. Of the three Australian species, one is also m 

 the 'Archipelago, the two others are endemic. The solitary erect ovules and seeds connect 

 this genns with Verbenaceee^ but the habit and corollas are those of FedalinecB, 



Leaves glabrons or nearly so, quite entire. Ovary usually S-celled. 



Fruit with a small terete or conical beak 1. /. grandiflora. 



Leaves pubescent underneath, the lower ones coarsely toothed. Ovary . . 



. usually 6 -celled. Fruit with a triangular truncate beak .... 2. J. imperatncis. 

 Plant densely villous. Lower leaves divided into 3 distinct segments. 



Ovary usually 4-celled. Fruit not beaked 3. /. Bagenia. 



1. J. grandiflora, R, Br. Prod. 520. Stems erect or diffuse (2 to 3 

 ft. high?), glabrous or sprinkled with a few minute hairs. Leaves petiolaie, 

 lanceolate or the lower ones ovate-lanceolate, all quite entire, 1\ to 3 in. 

 long, glabrous or minutely and sparingly pubescent underneath. ^ Pediceb 

 shorter than the petiole. Calyx-segments narrow, acuminate, about 2 lines lon?j 

 the upper one usually shorter. Corolla at least 1 in. lou^, pubescent outside, 

 the tube gibbous at the base on the upper side, gradually dilated upwards ; 

 lobes broad, the 4 upper ones nearly equal, the lower twice as long a^/ 

 broader than the others. Ovary in the flowers examined 8-celIed; stigmatic 

 lobes 4 (sometimes 3 according to Endlicher's figure). Fruit ovoid-globu- 

 lar, under \ in. diameter, very hard, villous with short soft hairs, armed witn 

 thick conical very unequal prickles, the persistent thickened base o*" ^^ 

 style forming a cylindrical or slightly conical beak, sometimes very short, 

 sometimes at least as long as the prickles. — Endl. Iconogr. t. 106. 



Queensland. Endeavour Straits, B. Brown ; Low Island, B. Brown^ Senne ; islan s 

 of Howick's group, F. Mueller; Three Isles and Lkard Islaud, M'GilUvrag; Pehcau an 

 Haggerstone Islands, A. Cunningham. 



.* 



