stamens of whorls I & II 1.3 - 1.8 mm long, their filaments pubescent ad and abaxially, anthers 
glabrous, stamens of whorl III 1.3 - 1.8 mm long, their filaments densely sericeous to glabrescent 
abaxially, glabrous adaxially, anthers glabrous, sporangia four in all stamens, glands ca. 0.5 mm 
long, rounded to elliptic, staminodes 0.9 - 1.1 mm long, filaments glabrous or sericeous apically 
on adaxial face, sericeous abaxially mainly on the margins, apex totally glabrous or pilose on the 
base abaxially, hypanthium 0.3 - 0.5 mm deep, sparsely tomentose or glabrous outside, glabrous 
or very sparsely pubescent inside, ovary and style glabrous; fruit 11.5 - 15.5 mm long, and 9 - 
11.5 mm wide, ellipsoid, seated on the persistent tepals, pedicel turbinate, reddish. 
This species grows mainly in oak and pine forest, but it has been found also in deciduous 
mixed forest, between 1000 - 1800 m, in a small area of northeastern Mexico. Flowers are 
produced during April and May (July), and fruits mature from July through September. 
Paratypes. MEXICO. Querétaro: Mpo. Jalpan, 4 -5 km S of La Parada, 1350 m, tree 6-8 
m (fl) B. Servin 168 (IEB, MO); Mpo. Jalpan, 3-4 km NE of El Saucito, 1150 m, tree 6-8 m (fr) 
B. Servin 220 (IEB, MO); Mpo. Jalpan, Cerro Los Fresnos, 2 km N of trail to El Carrizal, 1500 
m, tree 5 m (fr) C. Guzman 125 (IEB, MO); Mpo. Jalpan, 3-4 km N of La Parada, 1300 m, shrub 
2-3 m (fr) B. Servin 366 (IEB, MO); Mpo. Jalpan, Cerro El Pilon, S of La Parada, 1200 m, shrub 
2 m (fr) C. Guzman 58 (IEB). San Luis Potosi. Mpo. 82 km W of Ciudad Valles on highway to 
Rio Verde, 1400 m, tree 4 m (fr) P. Fryxell 3783 (ENCB, F, MO, NY); Mpo. 16-18 km NE of 
Ciudad del Maiz, 1400-1600 m, tree (fl) R. McVaugh 10443 (NY). Tamaulipas. Mpo. Gomez 
Farias, Rancho del Cielo, 1000 m, shrub 4 m (fl) A Gémez-Pompa 2039 (MEXU); Mpo. Gomez 
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