adaxially, sericeous abaxially, red trichomes present, anthers ca. 0.9 mm long, sericeo-villose at 
base adaxially, glabrous abaxially, sporangia four, upper ones latrorse, lower ones extrorse, 
glands ca. 0.6 mm long, above filament base, glabrous except for point of attachement adaxially, 
staminodes 1.2 - 1.4 mm long, filament sericeo-villose ad and abaxially, red trichomes abundant, 
head 0.8 - 1 mm long, triangular cordate, glabrous adaxially, sericeo-villose abaxially, sometimes 
sparsely so at upper half, hypanthium ca. 0.6 mm deep, glabrous or glabrescent outside, sericeous 
inside (sometimes in patches), red hairs abundant or scarse, pistil 2.1 - 2.3 mm long, glabrous, 
ovary | - 1.2 mm long, 0.9 - 1.1 mm wide; fruit 13.5 mm long, 10 mm wide, ellipsoid, cupule 9 - 
10 mm long, 7.5 mm wide above, 2.6 mm wide at the bottom, pedicel indistinct, tepals 
persistent. 
Flowers present in April and May. Ripe fruits present from August to September. No 
information about the type of vegetation where this species grows has been recorded, but it is 
suspected to be tropical rain forest, given the region where it has been collected. Altitudinal 
range is rather narrow, from 100 to 200 m. So far C. chavarrianum is known only from a very 
small area in central Costa Rica. 
This species is easily recognized on the basis of having leaves pinninerved lacking 
domatia, and fruits with persistent tepals. 
Specimens examined. COSTA RICA.Costa Rica. Alajuela. San Rafael de Guatuso, 
faldas del Volcan Tenorio, M. M. Chavarria et al. 236 (MO). Heredia. Finca La Selva, the OTS 
Field Station on the Rio Puerto Viejo just E of its juction with the Rio Sarapiqui, B. Hammel 
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