glabrous, ovary 1.1 - 1.3 mm long, 0.8 - 1.1 mm wide; fruit 11 - 21 mm long, 8 - 11 mm wide, 
ellipsoid, cupule 5 - 8 mm long, 5 - 7.5 mm wide, pedicel 3 - 5 mm long, 0.9 - 1.7 mm wide, 
tepals persistent. 
Flowering specimens have been recorded from February through April; ripe fruits can be 
found in December, January, February, July, and September. Natural vegetation where this 
species occurs include: cloud forest, premontane wet forest, and seconday forest, between 1000 - 
1500 (1700) m altitude. The distribution of C. costaricanum includes the mountains in central 
Nicaragua, and eastern mountain ranges in Costa Rica and adjacent Panama. 
C. costaricanum and C. neurophyllum are morphologically very close. Looking at sterile 
specimens or without fruits it is difficult to tell them apart. Here, the two species are kept distinct 
on the grounds of the correlation of few characters, namely the presence of persistent tepals, 
lowermost pair of secondaries not strongly raised, and domatia trichomes rather spreading to 
erect in C. costaricanum, and partially persistent tepals in fruit, lower most pair of secondary 
veins strongly raised, and domatia trichomes mostly parallel to leaf surface in C. neurophyllum. 
C. grisebachianum is barely different from C. costaricanum (see comment under that species). 
Specimens examined. COSTA RICA. Puntarenas. Las Alturas de Coto Brus, W. Burger 
et al 12190 (MO); Las Alturas de Coto Brus; ca. 20 km NE of San Vito, in forest along road 1-3 
km S of Las Alturas, B. Hammel et al. 14197 (MO); Buenos Aires, siguiendo filas en cuenca 
superior de Rio Cabagra., G. Herrera 3541 (MO); Cafias Gordas, H. Pittier 11107 (GH, BR, US 
(2sheets), NY); Foothills of the Cordillera de Talamanca, lower montane forest in the area of 
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