present, peduncle (0.3) 2 - 7 (8.5) cm long, puberulous, rachis puberulous to glabrescent, flower 
pedicel (3) 3.5 - 4.5 (5) mm long, glabrous; flowers , tepals 2.1 - 3.2 mm long, 1.9 - 2.4 mm 
wide, ovate or wide ovate, glabrous outside (inner ones with few hairs in lower altitude 
specimens), sericeous inside, sparsely so the outer ones (not so in low altitude specimens), 
stamens of whorls I & II 1.5 - 2.2 mm long, filaments sparsely sericeous adaxially along median 
section, glabrous abaxially (sericeo-villose ad and abaxially in low wltitude specimens), anthers 
0.9 - 1.5 mm long, glabrous ad and abaxially, sporangia four, introrse, stamens of whorl III 1.5 - 
2.2 mm long, sparsely sericeo-villose adaxially, glabrous abaxially (sericeo-villose ad and 
abaxially in low altitude specimens), anthers 0.7 - 1 mm long, glabrous ad and abaxially, 
sporangia four, upper ones latrorse, lower ones latrorse-extrorse, glands (0.5) 0.7 - 0.8 mm long, 
at filament base, glabrous or sparsely sericeous adaxially at point of attachement, staminodes 0.8 
- 1.5 mm long, filament sparse to densely sericeo-villose adaxially, glabrous abaxially or 
glabrescent (sericeo-villose ad and abaxially in low altitude specimens), head 0.5 -1.1 mm long, 
triangular cordate, glabrous ad and abaxially (sparsely sericeo-villose at base abaxially in low 
altitude specimens), hypanthium 0.5 mm deep, glabrous outside, glabrous inside, pistil 1.6 - 2.6 
mm long, ovary 0.9 - 1.1 mm long, 0.8 - 1 mm wide; fruit 18 - 20.5 mm long, 9 - 12.5 mm wide, 
ellipsoid, cupule 9.5 - 11.5 mm long, 6.5 - 8 mm wide, narrowing to a base 3.5 - 4.5 mm wide, 
pedicel indistinct, tepals partially persistent. 
Flowers are present from May to July; ripe fruits can be found in March and June. Habitat 
of this species includes primary forest and secondary vegetation, between 650 and 2500 m. The 
area of distribution goes from western and southern Costa Rica and western Panama along 
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