mountain ranges. 
As Burger mentioned in the original publication, C. hammelianum is morphologically 
similar to C. chavarrianum, and the only vegetative character that helps to distinguish that 
species from the latter one is the appearance of the lower leaf surface; light green and smooth in 
C. hammelianum, glaucous and papillose in C. chavarrianum. The best character is the 
persistence of tepals in the fruit; partially deciduous in the former, entire persistent in the latter. 
Unfortunately, contrary to what is found in other species, tepals in C. hammelianum fall off long 
after anthesis, making flowering material hard to separate. 
Type material of C. hammeliana (which is with flower buds only), for some reason 
mostly has anthers with aborted sporangia, looking rather like an Ocofea species. Fruiting 
material cited as paratypes seems to be conspecific with the type, but flowering material referred 
as paratype (from lower altitude) does not look similar to the type. However, there are no fruiting 
collections from the same lower altitude areas that could help to solve this problem. Thus, all the 
specimens that look similar to the material cited in the original publication is considered here 
under one name, although there might be two taxa involved. 
Specimens examined. COSTA RICA. Alajuela. Naranjo, Zona Protectora Chayote, 
Llevo Bonito, L. D. Gomez et al. 23977 (MO); Upala, Cordillera de Guanacaste, Bijagua, D. 
Penneys 414 (MO (7 dups)); Reserva Forestal de San Ramon, por sendero que sube a la fila hacia 
el SO de la estacion., G. Herrera 534 (MO); Along the Sarapiqui rd., about 1.5 km N of Finca 
Isla Bonita, G. Hartshorn 1456 (MO); San José to Vara Blanca on route 9 km 20, B. Hammel & 
M. Grayum 14091 (MO); Guanacaste. Parque Nacional Guanacaste, Est. Pitilla y Sendero El 
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