margins, glabrous or sparsely long sericeous abaxially, head 0.6 - 0.8 mm long, wide triangular 
in outline, acuminate, glabrous ad and abaxially, or sparsely sericeous at base abaxially, 
hypanthium (0.3) 0.4 - 0.6 mm deep, glabrous outside or sparsely pubescent, sericeous inside, 
sometimes sparsely so, red hairs sometimes present, pistil 1.9 - 2.3 mm long, glabrous, ovary | - 
1.2 mm long, 0.9 - 1.1 mm wide; fruit 9.5 - 13.5 mm long, 6 - 8.5 mm wide, ellipsoid, cupule 4 - 
8.2 mm long, 2 - 5.4 mm wide, tepals partially persistent, with an abscission line close to the 
middle, pedicel 1.5 - 2.8 mm long, 0.6 - 1.4 mm wide, distally swollen, turbinate. 
Most collections suggest this species flowers in early spring (March, April), but rarely 
also flowers in January. Mature fruits are found in May and July. The habitat where this species 
grows is in the contact zone of oak forests with semideciduous tropical forest or even in 
deciduous tropical forest, between (330) 500 - 1100 m altitude. Its geographical distribution 
extends on a wide area in the nortwestern Sierra Madre in Mexico, including the states of 
Chihuahua, Sonora, Durango, Nayarit and Jalisco. 
Vegetative morphology, particularly of leaves, is very variable in this species. However, 
the consistent features of tepals with an abscission line at or above their base, and persistent 
bases of tepals that tend to be erect and forming like a cup at the fruit base, make it a clear cut 
entity. 
Specimens examined. MEXICO. Chihuahua. Sierra Madre Occidental between Creek 
and La Bufa, canyon of Batopilas, on canyon floor near mouth of creek called Arroyo San 
Fernando, V. Siplivinsky et al. 4006 (NY); Ocampo River, H. LeSueur 1246 (F, MO); Temoris 
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