obviously does not belong to it, since that collection was referred as only with fruits in Nees 
description of Phoebe pachypoda. It is very likely that this specimen is partially the source of 
confusion found in Allen's (1945) concept of what was, in her view, Persea pachypoda. She 
describes the species as having "...densely tomentose..." flowers, which agrees with the flower 
bud slipped with type fragment at F, but wholly disagrees with the glabrous flowers of O. 
benthamiana type collection (Hartweg 84), cited in her work as synonymous with that species. 
Besides that, pubescence of that loose flower is made out of short and thick reddish-brown hairs, 
the kind of pubescence that Allen refers to as present in leaves and inflorescence axes, yet not 
found in the companion pickled leaves of Ehrenberg collection. The flower seems to belong to a 
Persea species (yet tepals seem to be about the same size in the bud), to which one is open to 
conjecture, but definitively does not belong to any American species of Cinnamomum. 
Fortunately, the specimen at GZU confirmed what Nees said in the description of P. pachypoda, 
and provided more information (fruit morphology, and more and bigger leaves) to conclude the 
connection of this and the Hartweg collection. The combination of leaves with lower surface 
velutinous, fruits with glabrous persistent tepals, and hypanthium glabrous inside, is not found in 
other species of the area. Furthermore, the now abundant collections of Cinnamomum from the 
area helped to solve the problem. There is flowering material from the area that matches Hartweg 
collection and, because of a wider representation of variation of morphology in the species, both 
the Hartweg and Ehrenberg collections are readily accomodated. The same is the case of Arséne 
2448 collection on which description of Phoebe arsenei was based. But in this case the inclusion 
of Arséne collection in Cinnamomum pachypodum is clearer. Pubescence and shape of leaves, 
flowers glabrous, and primary domatia usually inconspicuous, fit the Hartweg collection almost 
131 
