Flowers are present from March through May; fruits apparently take a long time for 
maturation, from August through January, but mainly in September - October. Tropical 
premontane forest is the natural type of vegetation where this species occurs, but it can be found 
also in secondary forest and pastures at an altitude between (800) 1100 - 1600 m. This species is 
known from central Costa Rica to western Panama. 
Paratypes. COSTA RICA. Guanacaste: Canton de Liberia, Parque Nacional Guanacaste, 
Cerro Cacao, 1100 m, (fr), C. Chavez 578 (CR, MO); La Cruz de Abangares, 1400 m, (fr), W. 
Haber & E. Bello 2861 (CR, MO); Canton de Abangares, upper San Gerardo Valley, 1500 m, 
(fr), W. Haber & W. Zuchowski 9534 (MO). Puntarenas: Reserva Biologica Monteverde, 
Arancibia de Miramar, 1600 m, (fr), E. Bello 1359 (CR, MO); Monteverde, Pacific slope, 1500 
m, (fr), W. Haber & W. Zuchowski 9529 (CR, MO); Reserva Biolégica Monteverde, Rio 
Veracruz, 1300 m, (fl), E. Bello 2776 (CR, MO); near the Reserve, 1520 m, (fl), V. Dryer 1335 
(MO); Monteverde, 1350 m, N. Wheelwright 126d (MO); Monteverde, 1350 m, (fr), N. 
Wheelwright 4b (MO); Monteverde, Pacific slope, 1450-1650 m, (fr), W. Burger & R. Baker 
9684 (MO): Monteverde, Pacific slope, 1500 m, (fr), W. Haber & E. Bello 3683 (F, MO). 
Limon: Canton de Limon, El Progreso, Cabecera del Rio Cariei, Valle de La Estrella, 1400 m, 
(fl), G. Herrera 2821 (MO). PANAMA. Chiriqui: Vicinity of Fortuna Dam, along trail to Rio 
Hornito, 1100 m, (fr), G. McPherson 1199] (MO); Panama: Parque Nacional Altos de 
Campana, | km E of rangers camp, 800-900 m, (fr), M. Correa & E. Montenegro 10793 (MO). 
The leaves without domatia or with domatia in just the basal pair of lateral veins, along 
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