present sometimes, sericeous abaxially, particularly on lower half, anthers ca. 0.8 mm long, 
sericeous on lower half adaxially, elsewhere glabrous, sporangia four, upper ones latrorse, lower 
ones extrorse-latrorse, glands ca. 0.5 mm long, on lower third of filament, ovate, sericeous 
adaxially on point of attachment, glabrous elsewhere, staminodes ca. 1.3 mm long, filaments 
densely sericeous ad and abaxially, red hairs present, head 0.8 mm long, cordate in outline, 
acuminate, glabrous adaxially, sericeous abaxially, hypanthium 0.5 mm deep, densely tomentose 
outside, red sericeous inside, pistil 2.2 - 2.4 mm long, ovary ca. | mm long, 1 mm wide; fruit 
11.5 - 18 mm long, 7 - 10 mm wide, cupule 5 - 8.5 mm long, 6.5 - 8 mm wide, sometimes 
narrowing gradually to a bottom ca. 1.5 mm wide, pedicel 1.5 - 3 mm long, 1.7 - 2.1 mm wide, 
or sometimes not distinct, tepals partially persistent. 
Flowers are present in February and March, and fruits in July and September. No specific 
type of vegetation is mentioned on specimens labels, but given the altitudinal range where this 
species occurs (1600 - 2100 m), it is likely the habitat to be cloud forest. The species is restricted 
to southeastern Costa Rica and western Panama, and apparently is not common. 
The presence of wavy to curly hairs on leaves, along with secondary domatia, and tepals 
partially persistent in fruit works well to separate this species from others in the area. Curly hairs 
and domatia suggest certain relationship to C. zapatae, but in the last species hairs tend to be 
persistent, and domatia are more abundant; besides this, tepals are persistent in the latter. 
Specimens examined. COSTA RICA. Puntarenas. Cerro Quijada de Diablo near the 
border with Panama, W. Burger et al. 12193 (F, MO); Buenos aires, Sabanas de Murur-Bisuk, 
173 
