paraguariensis have rather glabrous tepals outside, but there is a wide range of variation in this 
character among individuals across the study area. 
Plants with hairs spreading to erect on twigs, leaves with hairs spreading to erect on 
lower surface, secondary domatia sometimes present, and tepals pubescent outside correspond to 
types of Cinnamomum australe, Laurus triplinervis, and Phoebe brasiliensis. A slight variation 
to leaves without secondary domatia is represented by the type of Phoebe pickelii. 
Types of Cinnamomum chana, Phoebe filamentosa, and P. fruticosa represent specimens 
with hairs appressed to erect on twigs, leaves with appressed to spreading hairs on lower surface 
(very often glabrous with age), secondary domatia present, and flowers glabrous outside. The 
condition of leaves without secondary domatia in this group correspond to the types of 
Cinnamomum xinguense and Phoebe maynensis. An extreme of variation shown by plants with 
twigs and leaves (almost) glabrous, but with tepals (at least the inner ones) pubescent outside, is 
represented by the type of Cinnamomum portosecurianum. 
Specimens examined (selected). BELIZE. Cayo. Opposite Vaca, P. H. Gentle 2499 
(MO). Orange Walk. From mile 50, Northern Highway to new river toll bridge, J. D. Dwyer 
12755 (MO(3 sheets), ENCB). Stann Creek. Middlesex, P. H. Gentle 2941 (NY). Toledo. At 
base of hill beyond San Antonio, P.H. Gentle 7553 (MO); Hope creek, W.A. Schipp 281 (F); 
Hope Creek, P. H. Gentle 7937 (MO, F). COLOMBIA. Antioquia. Betania, Vereda Los Picos, 
Finca Los Manzanos, D. Sanchez et al. 1181 (MO); Hispania, 3.3 km de Hispania a la Vereda La 
Palmira, R. Fonnegra et al. 2482 (MO); Cocorna, H. Daniel 1628 (F); Cocorna, H. Daniel 1359 
(F); Amalfi, Valley of Rio Porce, 1-5 km SE of Puente Acacias (bridge over Rio Porce) towards 
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