widely ovate, pubescent outside, sericeous inside, some red hairs present, stamens of whorls I & 
II 1.6 - 1.8 mm long, filaments glabrous adaxially, glabrous or sparsely sericeous abaxially, 
anthers 0.7 - 1.1 mm long, glabrous ad and abaxially, or sparsely sericeous on base abaxially, 
sporangia four, introrse, stamens of whorl III 1.7 - 2 mm long, filaments sericeo-tomentose 
adaxially, red hairs sometimes present, sparsely long sericeous abaxially, particularly on lower 
half, anthers 0.7 - 1 mm long, sparsely sericeo-tomentose adaxially on lower half, glabrous 
abaxially, sporangia four, upper ones latrorse, lower ones extrorse, glands ca. 0.5 mm long, 
above filament base or higher, roundish, long sericeous adaxially, glabrous abaxially, staminodes 
0.9 - 1.1 mm long, filaments (red) sericeo-tomentose adaxially, sericeous abaxially, head 0.5 - 
0.7 mm long, wide cordate in outline, acuminate, glabrous adaxially, sparsely long sericeous on 
lower half abaxially, hypanthium 0.5 deep, pubescent outside, sericeous inside, red hairs 
abundant, pistil 2 - 2.3 mm long, glabrous, ovary 0.8 - 1.2 mm long, 0.9 - 1.2 mm wide; fruit 9 - 
15.5 mm long, 6 - 10 mm wide, cupule 5 - 9.5 mm long, 5 - 8.5 mm wide, narrowing to a bottom 
1.5 - 2.5 mm wide, pedicel not distinct, tepals persistent. 
Flowers are present in January, April, and May. Fruits are ripe in December, January, and 
February. The habitats of this species are mainly the cloud forests and pine-oak forests that grow 
along the mountain ranges that run across southern Mexico and adyacent Guatemala, between 
(1300) 1600 - 2450 (2750) m of altitude. 
Paratypes. MEXICO. Guerrero: Mpo. Chichihualco (Leonardo Bravo), 3.5 km NE of 
Cruz de Ocote, 2100 m, (fr), F. Lorea 5489 (FCME, MO); Mpo. Chilpancingo, 2 km E of 
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