although Hortonia recently has been reported as sometimes having toothed leaves, no record of 
lobed leaves exists for this genus or any other closely related to it. Thus, entire leaves are 
considered the ancestral state. 0, entire leaves; 1, lobed leaves. 
4. Persistence of leaves. At first this character was considered the product of 
enviromental adaptation, but the fact that several taxa from temperate regions do not present 
deciduous leaves suggests the likelihood of this being a feature developed in one or few phyletic 
lines. Taxa with perennial leaves (with this is meant that leaves and flowers are present at the 
same time) outnumber those with deciduous leaves. The three species of Hortonia have perennial 
leaves. 0, leaves perennial; 1, leaves deciduous. 
5. Domatia in leaves. These specialized areas of the lamina are found only in some genera 
of the Lauraceae. They are not present in Hortonia. 0, domatia absent; 1, domatia present. 
6. Inflorescence protection. Based on figs. 2, 3 and 4 from Qiu et al. (1993), this character 
can be polarized following the method proposed by Maddison et al. (1984). This indicates that 
inflorescences without bracts are the ancestral condition for this character. 0, bracts absent; 1, bracts 
present. 
7. Inflorescence architecture. This character is rather difficult to polarize. Cymose- 
paniculate inflorescences, as well as racemose ones, are found both in the Monimiaceae and the 
Lauraceae. Inflorescences in two of the three species of Hortonia (H. angustifolia and H. 
floribunda), present at least partially cymose-paniculate inflorescences. Based on this evidence 
cymose-paniculate architecture is considered plesiomorphic. 0, cymose-paniculate; 1, botryoid or 
racemose; 2, umbellate. 
8. Floral merosity. As was mentioned above, recent surveys about the phylogenetic 
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