154 
The genus Souliea, separated from Isopyrum by Franchet, 
forms the bridge between Paraquilegia and the Cimicifugeae, and 
of the latter an extreme reduction may be the remarkable genus 
Achlys in Berberidaceae, which has the inflorescence of Cumict- 
fugeae, but has no perianth. Then again the genus Enemion 
seems to be a direct offshoot from Paraquilegia, the divergence 
here being the entire reduction of petals and the tendency to an 
umbellate arrangement of the flowers. The remarkably close 
affinity of two species of this genus, H. Raddeanum from Man- 
churia, and £. Hallii from Oregon, is worthy of note. 
Asteropyrum (fig. 1), a genus described below for the first 
‘ time, is not shown in the phylogenetic table above because its 
oie. postion seems to be in the neighbourhood of Caltha and 
ius 
are much indebted to Professor Bayley Balfour and Dr. 
A. B. Rendle for allowing us to examine the material in the 
herbaria under their charge, to Miss E. Milner for general assis- 
tance, and to Miss D. M. Rolfe for help with the text figures. 
Key to the genera separated from Lsopyrum. 
Flowers solitary or rarely subumbellate; car- 
pels more 1: 
Staminodes absent : 
Carpels several, always more than 2: 
etals present, sometimes small and 
much modifie 
Leaves peltate ‘carpels spreading 
stellately in fruit; petals stipi- 
tate and indusiform (see fig. 1) 1. Asteropyrum 
Leaves not peltate, usually much 
divided : 
Caespitose perennials; carpels 
usually about 5, mostly erect 
in fruit; petals not —— 
at the base Oe 2. Paraquilegia 
(ge 
Annuals; carpels about 12 or 
more; petals tubular at the 
base... vais ee ... 3, Leptopyrum, 
Reichb. 
ee —— inflorescence often um- 
4. Enemion, Raf. 
ees ah 2 ( —- 3), divaricate i in n fruit; 
per 5. Isopyr hes 
oe 8 gg mem mbranous and flat within ae 
the —e stamens; ares saccate at 
the bas 6. Semiaquilegia, 
Mak. 
Flowers several in short racemes ; are 
vaginate at the base; carpel 1 2. Te Souliea, 
neh, 
