164 OENOTHERA LAMARCKIANA MUT. VELUTINA. 



If we now try to resume this description in such terms as to 

 distinguish a number of probable units, the combination of which 

 might constitute the type of velutina, I might propose the following: 

 (1) slender stature; (2) long internodes of the flower spike; (3) leaves 

 narrow, folded longitudinally and smooth, that is, without bubbles; 

 (4) flowers cup-shaped; (5) hairiness of all organs; (6) abundance 

 of red color in the younger parts. It is obvious, however, that some 

 of these points may go together and depend upon one unit, but on 

 the other hand it must be conceded that this list may be far from 

 complete. Most of these assumed units are recessive to the corres- 

 ponding qualities of 0. Lamarckiana, but the smoothness is dominant 

 over the bubbles, which are evidently due to a lack of growth parallel 

 to the surface of the blade of the leaf. This enables us to separate 

 smoothness from the larger part of the other characters by means 

 of crosses, as we shall see when dealing with the hybrids. 



I shall now describe the origin of 0. blandina and the pedigree 

 of the race derived from it. As a matter of fact, this mutant did 

 not arise directly from 0. Lamarckiana, but through another mutant 

 race as an intermediate. This was the fertile race of 0. lata issued 

 from an original fecundation of my normal 0. lata by means of the 

 pollen of 0. semilata. This race is described in my book Gruppen- 

 weise Artbildung on pp. 256, 257. In the fourth generation of this 

 race, issued from the seeds of 1904 sown in 1905, 1907, and 1908, 

 three specimens of 0. blandina were observed, two in 1907 and one 

 in 1908. Besides these, the cultures consisted of specimens of 0. 

 lata and 0. Lamarckiana in the usual proportions, and of other 

 mutants such as 0. nanella, 0. oblonga, and 0. scintillans. The size 

 of the cultures, however, was too small to calculate percentage figures. 



The full name of the new type, therefore, would be 0. Lamarckiana 

 (mut. 1888 lata x mut. 1895 semilata) mut. 1907-8 blandina, 

 leaving out the fecundation of the lata specimens of the pedigree 

 by the pollen of 0. Lamarckiana. 



The second generation was derived separately from one mutant 

 of 1907 and from that of 1908, both after guarded and artificial 

 pollination. The seeds of the first mutant were sown in 1913 and 

 gave a culture of 25 flowering individuals and 45 others which were 

 pulled up shortly before flowering. Both groups were uniform, apart 

 from the variation in the green color already mentioned. They 

 were fully like the simultaneous culture of the third generation of 

 the other strain, and for this reason this first line has not been 

 continued. 



