OENOTHERA GRANDIFLORA AIT. 221 



but also one of the mutants of each of them, 0. Lamarckiana nanella 

 and 0. grandiflora lorea. Their special characters were latent in the 

 first generation, and the results of their crosses were identical with 

 those of the species. They simply give a confirmation of the main 

 results. This consisted in the appearance of three types, which it 

 seems desirable to distinguish by special names. I shall call these 

 triple hybrids ovata, lutea, and brunnea, in connection with their most 

 striking features. The mutant previously mentioned, for which I 

 have not succeeded in studying the identity with one of the mutants 

 of the parents, I shall call, for convenience, contraria. It seems 

 destined to play only a subordinate role in the discussion con- 

 cerning the splitting which produces the triple hybrids. 



0. hybr. ovata is seen almost always in the largest numbers. It 

 is stout and richly branched, with broad leaves of pure green and 

 dense spikes of large flowers. In spring it makes a large rosette, 

 like those of 0. Lamarckiana, and from its center produces a vigorous 

 stem in June. The foliage is that of 0. grandiflora, and the leaves 

 in the middle part of the stem are hairy, show some bubbles, and 

 measure approximately 4 x 17 cm. The combination of this foliage 

 with the thick stems and branches of 0. Lamarckiana constitutes 

 the most striking mark; the hybrid is obviously intermediate between 

 its parents. The flower buds resemble those of Lamarckiana and have 

 a tinge of brown, which is subjected to some amount of fluctuating 

 variability. The flowers are large and the petals cover one another 

 by the margins, but this mark is also variable, and sometimes the 

 petals are seen to have a wedge-shaped base. The fruit are stout 

 and of an intermediate form. 



0. hybr. lutea differs little from ovata in form, but its color is 

 strikingly yellowish instead of pure green. This is seen in the foliage 

 and very evident in the flower buds, but may disappear when 

 the season advances. The leaves are almost as large. I measured 

 strictly comparable instances for comparison with the figures just 

 given and found 4.5 x 14 cm., that is, a little broader and shorter 

 than ovata. The spike is destitute of red color, or almost so, in all 

 its parts, and this is often the most striking mark. The flower buds 

 are pale and stand off at wide angles from the axis. The flowers 

 are large and the petals have usually a broad base, as in 0. Lamarcki- 

 ana. The stems are not as stout nor as richly branched as in ovata, 

 but this is probably due to the less amount of chlorophyll. The 

 fruits are like those of the other type. 



0. hybr. brunnea is a very striking form, especially when culti- 



