1913] Life-History of a Bee- Fly. 223 



mixed pine and cottonwood areas but none of these have been 

 found with parasites. The large pit and goose-neck burrow- 

 probably prevent this species from serving as a host (Shelford, 

 '08). Very few parasitized Cicindela Leconti larvae have been 

 taken here though the host is only a little less abundant than 

 in the oak* area. It is on the margins of the depressions in the 

 pine areas that the larvae of C. tranqueharica Hbst. are numerous 

 (Shelford '07) but none of those of this species were found 

 parasitized, though the number of larvae dug was great. 



Spogostylitm anale and its host species are most abundant 

 in the early stages of the black oak forest where cacti occur 

 (Fig. 3). The exact landscape aspect is significant only as an 

 index of the physicial conditions. The evaporation in these 

 open oak forests is about one half that of the cottonwood 

 area and less than that of the pine area. The available soil 

 moisture is less (Shelford, '12). • 



2. Geographic Distribution. 



American dipterologists have kindly supplied me with 

 data on the distribution of Spogostylum anale as follows: 



Prof. D. W. Coquillett: Sandy Hook, N. J.; Indiana; 

 Carbondale, 111.; Mississippi; British Columbia; Washington; 

 Mesilla, N. M.; St. Louis, Mo.; Shreveport, La.; Georgia; 

 Enterprise, Fla. ; Cambridge, Mass. 



Dr. J. S. Hine: Mission, British Columbia; Onaga, Kansas; 

 Akron, Ohio; Cincinnati, Ohio. 



Dr. C. T. Brues: Douglas Co., Kansas; Crete, Nebr. ; 

 Austin, Texas. 



Dr. C. F. Adams: Jackson Co., Mo.; Lawrence, Kan.; 

 Clark Co., Kan.; Fayetteville, Ark. 



Prof. E. P. Felt: South Britian, Conn.; Albany, N. Y. 



Prof. J. H. Comstock: Manlius, N. Y. 



Dr. A. L. Meander: Galveston, Texas; Golden, Colo. 



Map (Fig. 17) shows the distribution area of the host 

 (A) generously represented by connecting the more remote 

 localities along nearly direct lines where suitable habitats are 

 known to occur. The distribution of the parasite is wider 

 than that of the host species including all varieties extending 

 into Mexico and along the Pacific coast. Accordingly the 

 parasite must use another host. Several other species of the 

 tiger beetles may no doubt serve this purpose. 



