i93 2 Chromosome numbers in Hemerocallis, with reference to triploidy 253 



slightly different in coloring, and the flowering begins somewhat later 

 in the summer. There is much abortion of pollen, and there is complete 

 self -incompatibility, but fruits and a limited number of viable seeds per 

 capsule are readily obtained from certain compatible cross-pollinations. 

 The chromosome counts in the root-tips of plants of this clon are 

 most frequently 33 and the clon is evidently a triploid. 



The double-flowered fulvous daylilies. The varietal names " Kwan- 

 so" and "flore-pleno" have been given to certain double-flowered 

 daylilies that are very similar to the Europa Daylily in habits of growth 

 and flower colors. These are propagated exclusively by division and 

 have evidently long existed in China and Japan where they appear to 

 have escaped widely from cultivation. Plants of the double-flowered 

 type were not brought to Europe and America until about 1860 (see 

 Stout, 1929 a). Plants have been received at The New York Botanical 

 Garden under the names " Kwanso " and " Flore Pleno " from various 

 nurserymen and from various botanical gardens. The counts of chro- 

 mosome numbers in root-tips of these have, as a rule, been 33, as 

 reported for the plants of "Kwanso" studied by Takenaka. A study 

 of the cytology of sporogenesis in " Hemerocallis fulva fl. ploeno" by 

 Sienicka (1929) reports " 16 gemini " in stages of diakinesis and many 

 irregularities in the later stages of meiosis and spore formation. It 

 seems very probable that the plant studied by Sienicka is a member of 

 one of the triploid clons. 



A double-flowered fulvous daylily having variegated foliage was 

 mentioned in the Gardeners' Chronicle in 1864, under the name "H. 

 Kwanso foliis variegatis". Possibly the original plants of the so- 

 called " H. fulva Kwanso " brought by von Siebold directly from Japan 

 to Europe shortly before 1864 were variegated, for the variegation is 

 evidently of the chimeral type and the plants frequently produce all- 

 green branches. The root-tips of several plants of the variegated type 

 have been examined and in these the number of chromosomes is 33. 



Seedlings of Crosses between Diploids and Triploids 



The triploid clons Europa and Maculata are evidently fully self- 

 and cross-incompatible but they have been used both as seed and as 

 pollen parents in crosses with certain diploids. 



As a seed parent the Europa Daylily crossed with diploids has 

 given for the plants thus far studied— 



Cytologia3. 1932 18 



