303 



developed, and much exceeds the calyx, and the limb is almost 

 actinomorphic ; whereas in Markhamia the cylindric part of the 

 tube is very short and concealed in the calyx, only the upper part 

 of the funnel being visible, and the limb is conspicuously 

 bilabiate. In addition, Dolicliandrone has pure white, fragrant 

 flowers which are open during the night, whilst those of 

 Markhamia are yellow (rarely pink or lilac), or have a yellowish 

 tube, spotted with purple, and brown-purple lobes, and expand 

 in the daytime. Roxburgh says of D. crispa : "the large pure 

 white fragrant flowers . . . expand in the evening and drop in 

 the morning."* Wight states that those of D. arcuata usually 

 expand in the evening and drop off in the morning, t and Ridley 

 describes the flower of D. Rheediit as u white, with a long tube, 

 and a spreading circle of petals, and very fragrant, scenting the 

 air deliciously in the early morning. The flowers are nocturnal, 

 apparently opening after dark and commencing to fall by seven 

 o'clock in the morning." The nocturnal expansion, white 

 colour and powerful scent of the flowers, and the long, slender 

 corolla-tube all point to the pollination of Dolicliandrone by 

 moths. ~No observations as to the pollination appear to have been 

 published, and the genus is not mentioned in Knuth's Handbuch 

 der Bliitenbiologie. 



It seems desirable to give an enumeration of the species 

 referred to Dolicliandrone and Markhamia respectively, as the 

 synonymy is rather involved, and some of the species are not veTy 

 well known. 



Dolichandrone, Seem, in Ann. Mag. Nat. Hist. ser. 3, vol. x. 

 p. 31 (1862); in Journ. Bot. 1863, p. 226; et I.e. 1870, p. 379; 

 Benth. et Hook. f. Gen. Plant, vol. ii. p. 1046, partim (section- 

 ibus Markhamia et Muenteria exclusis) ; R. Schum. in Engl, et 

 Prantl, Nat. Pflanzenfam. vol. iv. 3 B, p. 75. — Spathodea, E. Br. 

 Prodr. p. 471; Bureau, Monogr. Bignon. p. 50; non Beauv. 

 Dolichandra, sect. Dolicliandrone, Fenzl in Denkschr. Baier. 

 Bot. Ges. Kegensburg, vol. iii. p. 265 (1841). 



A. Species australianae. 



1. D. alternifolia, Seem, in Journ. Bot, 1870, p. 340; Bailey, 

 Class. Index PI. Queensl. p. 29 (1883); Queensl. FL p. 1135. 

 D. heterophylla, F. Muell. Fragm. vol. iv. p. 149, partim; Seem, 

 in Journ. Bot. 1870, p. 382, partim. Spathodea alternifolia, 

 R. Br. Prodr. p. 472 (1810); DC. Prodr. vol. ix. p. 209; Bentli. 

 Fl. Austral, vol. iv. p. 538. 



Distrib. Queensland. 



F. Mueller and Seemann considered D. alternifolia as a mere 

 form of D. heterophylla, but Bentham, followed by F. M. Bailey, 

 treated it as a distinct species; and this seems to be the best 

 course to adopt in the present state of our knowledge. The 

 Queensland specimens quoted by Bentham under D. heterophylla 



* Fl. Ind. vol. iii. p. 104. 



t Wight Ic. vol. iv. pt. 2, p. P. 



2 Journ. As. Soc. Straits, no. 59, p. 40. 



