154 CRUCIFERiE. Dentaria. 



D. maxima, Xutt. Tubers near the surface aud stems reclined in fruit : leaves two or 

 three, alternate, 3-foliolate ; leaflets ovate or oblong-ovate, coarsely toothed and somewhat 

 cleft or lobed, 1 or 2 inches long: pods as in D. laciniuta ; seeds round-oblong; cotyledons 

 unequal, the smaller cuneate-oblong, half as wide as the larger ; radicle acute and cur^^ed, 

 cleft to the middle. — Gen. ii. 66; Gray, Gen. 111. i. 138, t. 56. D. laciniuta, var. 5, Torr. & 

 Gray, Fl. i. 86. Cardamine maxima, Wood, 1. c. — Vermont, Morgan, to Penn.sylvania and 

 Western New York ; said to bloom two weeks later than D. laciniata. Nuttall's original 

 specimens from Pennsylvania and W. New York are described as two feet tall and with five 

 to seven leaves. Nothing corresponding to tliis appears to have been found since. The 

 single small specimen, so named by Nuttall in the herb. Brit. Mus., from Pennsylvania, 

 has a pair of separate teniate leaves and probably belongs to this species or possibly to 

 D. diphylla. 



-H- -H- Western species. 



D.* macrocarpa, Nutt. Glabrous or slightly pubescent : stems simple, 4 to 15 inches high ; 

 joints of the rootstock about an inch long- leaves 1 to 3, approximate, shortly petiolate, 

 palmately or pinnately 3-5-parted or -divided, the segments linear to oblong, entire, obtuse 

 or acute, h, to 2 inches long ; basal leaves sometimes merely lobed or cleft ; the leaflets 

 sessile or petiolulate, often 3-5-lobed or -toothed: raceme usually nearly sessile: flowers 

 purple or rose-color: pods one or two inches long (including the style, usually 3 lines long) 

 and a line broad ; stigma capitate and entire ; seeds obloug ; cotyledons somewhat unequal ; 

 the oblique radicle cleft to the base. — Nutt. in Torr. & Gray, Fl. i. 88 ; Brew. & Wats. 

 Bot. Calif, i. 30. D. tenella, Brew. & Wats. 1. c. as to pi. Calif. ; AVats. ms. of present work, 

 not Pursh. D. ;/emmata, Wats, as to pi. of Howell, Pacif. Coast PI. 1887 (not as to type 

 which was later identified with D, tenella), Cardamine NuttalUi, Greene, Bull. Calif. 

 Acad. Sci. ii. 389. C. geinniata, Greene, Pittonia, i. 162. — N. California (Plumas and 

 Siskiyou Counties) to Brit. Columbia, LyaUA 



Var.* pulcherrima, Bobinson, n. var. Flowers larger than in the type: petals 6 to 

 8 lines long, 4 to 5 lines broad. — Cardamine pulcherritiia, Greene, Erythea, i. 148. — Hosier, 

 Oregon, T. Howell. Very nearly related species, if distinct at all, are the following: 

 CardAmine sinuAta, Greene, 1. c, with suborbicular sinuate-dentate radical leaves and roots 

 .said to be tuberous, from Crescent City, Calif., T.Howell, and (?) Cow Creek Mts., Oregon, 

 Henderson; also C. quercetorum, Howell, Erythea, iii. 33, with radical leaves 3-foliolate; 

 leaflets ovate to elliptic-oblong, dentate, from Silverton, Oregon, T. Howell. In their flowers, 

 young fruit (so far as known), canline foliage, and general habit, these plants show such 

 a close resemblance to each other aud to more robust forms of D. macrocarpa, that the 

 specific distinctions, derived chiefly from the subdivision of the radical leaves (in this genus 

 notably inconstant), appear very doubtful. Good specimens of the roots (not at hand) may 

 furnish better distinctions. 



D.* tenella, Pursh. Rootstock bearing small irregular tubers : basal leaves simple, round- 

 cordate, coarsely crenate or sinuate, one or two inches broad ; the petiole bearing usually 

 several clusters of bulblets : stem 6 to 12 inches high, with one or two nearly sessile 3-folio- 

 late leaves, sometimes bulbiferous in tlie axils ; leaflets linear-oblong or linear, obtuse, entire, 

 I to 2i inches lojig: raceme sessile or shortly pedunculate : flowers rose-color: pods an inch 

 long and a line wide, with a slender style tipped with a broad distinctly lobed .stigma. — 

 Fl. ii. 439 ; Torr. & Gray, Fl. i. 87. D. Unuifolia, Hook. Fl. Bor.-Am. i. 46, not Ledeb. — 

 Banks of the Columbia, Lewis ; Washington, Klikitat Co., Suksdorf, Upper Nesqually Val., 

 Allen. 



D. Californica, Nutt. Tubers of the submoniliform rootstock mostly small : stem ^ to 2 

 feet high, rather stout, simple or branched above, glabrous or slightly pubescent : foliage 

 very variable ; basal leaves entire or 3-foliolate ; tlie leaflets petiolulate, suborbicular, cune- 

 ate to subcordate at base, sinuate or coarsely tootlied ; cauline 2 to 4, mostly shortly petio- 

 late and above the middle of the stem, 3- or pinnately 5-foliolate, rarely simple or lobed ; 



1 The treatment of D. macrocarpa and the following species has been revised in the light of more 

 copious material. Nuttall's .species was unfortunately cliaracterized as liaving 3-foIiate radical leaves 

 Avith "reniform " leaflets. It is stated that the .species was founded u]ion a single specimen, and a 

 l)lant, so labelled by Nuttall himself, is now in herb, Brit. Mus., and is (ace to Dr. Watson) of the 

 species here described. 



