434 SAPINDACEiE. Staphylea. 



9. HYPELATE. Sepals 5, subglabrate, rounded, imbricated. Petals 5, unappendaged, 

 sessile. Ovary 3-celled ; ovules 2 in each cell, superposed, the upper ascending, the lower 

 somewhat pendulous. Fruit 1 -celled, 1 -seeded ; embryo with thin somewhat crumpled 

 cotyledons incumbent upon a long radicle. 



* * * * Fruit a coriaceous 3-valved loculicidal capsule : tree or erect shrub with pinnate 

 leaves and showy zygomorphous flowers : ovary and capsule stipitate. 



1 0. UNGNADIA. Calyx deeply 5-parted ; segments imbricated in the bud. Petals mostly 

 4, obovate-spatulate, uuguiculate, each with a bushy crest at the summit of the woolly- 

 pubescent claw. Disk thin, ol)liquely developed, bearing the stamens and closely envelojiing 

 and adnate to the stipe-like base of the ovary. Stamens mostly 8, unequal, much exserted 

 in the cf flowers, in the ? shorter than or about equalling the petals. Seeds by abortion 

 solitary in the cells, brown, smooth and shining, carunculate-appendaged at the broad 

 hilum. 



Suborder V. HIPPOCASTANE-^. Flowers irregular, polygamous, showy. 

 Sepals or calyx-lobes 5. Petals 5 (or 4), uuguiculate. Leaves opposite, palmately 

 5-9-foliolate. 



11. ^SCULUS. Calyx cup-shaped or tubular, usually unequal and gibbous. Petals hy- 

 pogynous, irregular, unappendaged. Stamens 5 to 8, usually 7, declined, exserted (at least 

 in sterile flowers). Ovary 3-celled ; cells each with a pair of superposed ovules. Fruit cap- 

 sular, loculicidal ; cells by abortion mostly only 1 or 2, and 1-seeded ; the valves thick, cori- 

 aceous, often spincscent or spiny. Seeds large, brown, smooth and shining, with large 

 dull white hilum. 



1. STAPHYLEA, L. Bladder-nut. (^racjivXy], a bunch of grapes, re- 

 ferring to the clustered flowers and fruit. The name as first applied by Tourne- 

 fort was Stapkylodendron.) — Shrubs with opposite stipulate pinnate 3-7-foliolate 

 leaves, nodding racemosely or cymosely paniculate white flowers, and strongly 

 inflated 3 (or rarely 2)-lobed capsules. — Spec. i. 270; Lam. 111. t. 210; Schk. 

 Handb. t. 84; Deless. Ic. iii, t. 51 ; Gray, Gen. 111. ii. 191, t. 172 ; Benth. & 

 Hook. Gen. i. 412 ; Zabel, Gartenfl. xxxvii. 498-504, 527-531 ; Pax in Engl. & 

 Prantl, Nat. Pflanzenf. iii. Ab. 5, 260. Staphylodendron, Tourn. Inst. 616, t. 

 386. Staphyllodendron, Scop. Fl. Carn. ed. 2, i. 223. Bumalda, Thunb. Fl. 

 Jap. 8. — A small genus of some eight species ; one European, three or four Asiatic 

 (often cult, for ornament and frequently hybridized), one Mexican, and the 

 following. 



S. trifolia, L. A neat shrub, 6 to 12 feet high, with green branchlets and 3-foliolate pubes- 

 cent or glabrate leaves : leaflets ovate, sharply acuminate, finely serrate, paler beneath : 

 sepals oblong, greenish white : petals a little longer, sub-unguiculate, with broad pubescent 

 claws : stamens and pistil scarcely or not at all exserted : filaments pubescent : the 3-horned 

 bladdery capsules 1^ to 2 inches long. — Spec. i. 270 ; Torr. Fl. N. Y. i. 139, t. 19 ; Gray, Gen. 

 111. ii. 192, t. 172; Chapm. Fl. 77. S. trifoliata, Schmidt, Arb. ii. t. 81. Staphylodendron 

 trifoliatum, Mcench, Meth. 65. — Rich moist soil. Lower Canada to Minnesota and southward 

 to North Carolina, Tennessee, and Missouri, Bush ; common ; fl. May ; fr. June ; autumnal 

 flowers and fruit also occur. Zabel's vars. typica & panel flora (the latter based apparently 

 upon cult, pi.) are not clearly di.stinguishable in the specimens at hand. 



S. Bolanderi, Gray. Leaves glabrous, 3-foliolate ; leaflets broadly ovate-oblong or suborbicu- 

 lar, scarcely acuminate, finely serrate, paler beneath, thickish (at least of firmer texture than 

 in preceding) : stamens and pistil much exserted nearly twice the length of the floral en- 

 velopes ; filaments not always (as originally described) glabrous : fruit 15 to 18 lines in length, 

 sharply 3-horned. — Proc. Am. Acad. x. 69; Brew. & Wats. Bot. Calif, i. 108; Sargent, 

 Gard. & For. ii. 544, t. 142. — California, on McCloud's Fork, Shasta Co., Bolander, Lem- 

 mon ; Fresno Co., Parry ; also near Sequioia Mills (ace. to Braudegee) ; fl. April; fr. July; 

 rare. 



