68 H. H. SWINNERTON, 
though it ends in a piece of cartilage and abuts against the clavicle 
at its upper end, has lost all connection with the dorsal edge of the 
girdle. Owing to the peculiar shape of the whole, the glenoid border 
has a shape intermediate between a semicircle and a right angle. Of 
the four radials (pterygia, pterygiophores, ra. 1—4) the two preaxial 
ones are slender, the others stout, and the fourth wholly cartilaginous. 
Resting on the distal ends of the two postaxial radials is a carti- 
laginous nodule, the sole representative of the familiar series which 
in other fishes give attachment to the fin rays, and are usually as 
numerous as these. 
In Galaxias (Fig. P) the posttemporal is simple and is attached 
to the epiotic. A postclavicle (p. cl) is present. Unlike Cromeria 
it has all the elements of its primary girdle lying in the same plane 
-— nearly vertical. The ossifications are so extensive that most of the 
cartilage which remains is confined to the narrow strip between the 
coracoid and scapula. Thus the histological condition of the girdles 
of these two fishes reverses that of their crania. Here again the 
coracoid meets its fellow; but, owing probably to the flattening of the 
posterior region of the girdle, the mesocoracoid has disappeared. The 
glenoid border is straight; the four radials are ossified, and the 
series of cartilaginous nodules at their extremities is complete. 
Apart from the elongation of the so called basipterygia in the 
pelvic girdle of Cromeria, and the presence of three cartilaginous 
radials in both types, there is nothing to be said about the skeleton 
of the hind-limbs. 
Remarks. 
The above account of the osteology of Cromeria shows that this 
is an extremely specialized type. The features which more particularly 
point to this are: the very complete ossification of the cranium, 
the enormous frontals and supraoccipitals and diminutive parietals, 
the large fontanelle in the cranial roof caused by the unusual size of 
the brain, the fusion of the mesethmoid with the nasals and suborbitals, 
the reduction of the ethmoid region, the fusion of the vomer and 
parasphenoid, the absence of symplectic!), the fenestration of the 
joints in the branchial skeleton, the lineal arrangement of the 
pharyngobranchials, and the attachment of the posttemporal to the 
1) This feature is shared by Cromeria with the Apodes and some 
Symbranchia; v. Jorpan, D. S.,, and Evmrmann, B. W., 1. c. 
