78 H. HEATIT and M. H. SPAULDING, 



Looking at the reproductive cap from above it appears to be 

 composed of tubes coiled irregularly into a circular disc at the right 

 (left in Fig. lU) side of which are situated the tumid lips of the ovi- 

 duct and albuminiferous and muciparous glands. The more detailed 

 relationships of these glands are shown in the diagrammatic section 

 of the visceral mass. Here it is seen that generally speaking the 

 upper layers of this glandular cap are composed of albuminiferous 

 cells while the muciparous ones constitute the lower and usually 

 larger canals. In each set of glands the walls are thrown into folds 

 of varying size which divide the internal cavity into spaces that are 

 proportionately variable. In every case these pockets communicate 

 with a larger more central cavity. The latter space in the case of 

 the albuminiferous (upper) gland by a somewhat tortuous course 

 opens to the exterior beside the external opening of the oviduct. On 

 the other hand the muciparous gland has no direct connection with 

 the exterior, its means of communication being by ducts with the 

 oviduct and by a connection (indicated by an arrow, Fig. 12) between 

 its cavity and the lumen of the albuminiferous gland. 



In every one of upwards of a dozen individuals carefully examined 

 the seminal receptacle contained large quantities of spermatozoa. In 

 fully three times this number of specimens the gonad contained ova 

 in all stages of development but no trace of sperm cells. This appears 

 to the case with all the species of the genus. Peck speaking of 

 C. calceola says: "Not only is the generative gland, in all these speci- 

 mens, in a state of female activity, but I have been quite unable to 

 find any penis to indicate that they ever function as males, or at 

 least so as to effect a copulation among themselves." A ciliated band 

 of cells is described by Peck as "leading away from the external 

 opening of the genital organs, but it becomes less and less con- 

 spicuous in its anterior course until it is lost near the opening into 

 the inner palliai cavity; but it does not lead into any groove or 

 cavity, nor is there any indication of its relation to any penis". In 

 C. spectahilis even this ciliated tract is absent and there is otherwise 

 no data to enable one to decide if the penis has entirely disappeared; 

 or whether the species are true hermaphrodites; or whether they may 

 be progressively hermaphrodite ; or finally if the males may not as yet 

 be undiscovered or possibly described as another genus. 



