The brain of Acipenser. 163 
of communis components. STRONG (95) describes the innervation 
of the end buds in the tadpole, where they are confined to the mouth 
and pharyngeal cavities. The rami innervating the regions in which 
end buds are found are composed chiefly or wholly of communis 
fibres and it can scarcely be doubted that the buds are innervated 
by these. GORONOWITSCH (96) states that communis components 
enter the N. weberi (N. lateralis accessorius) and the R. ophthal- 
micus profundus. ALLIS (97) finds the following rami apparently 
innervating end buds in Amia: R. ophthalmicus V, R. maxillaris V, 
accessory rami of V, r.ghs and r.ghi of V, and rami of IX and X. The 
first two of these rami he traced directly to end buds and he describes 
them as made up of communis fibres. The weight of ALLIS’ evidence 
is very much weakened by a curious error in analyzing the ganglionic 
complex. The posterior dorsal portion of his anterior root in Ama is 
undoubtedly the dorsal V (Va of STRONG). According to ALLIS, 
the fibres of this root passed into the anterior portion of the 
ganglionic complex, into the portion from which the rami ophthal- 
micus and maxillaris arose, and “disappeared gradually as they ap- 
proached the truncus maxillaris” (p. 594). He nowhere mentions 
these fibres in any nerve rami, but derives the two important rami 
of the V above mentioned wholly from the communis system. All 
other authors have described these rami as made up largely or ex- 
clusively of fibres from the root which Auuıs allows to be lost in 
the ganglion, and general cutaneous innervation requires the presence 
of this component. If the rami in question do contain communis 
fibres as ALLIS states, it is undoubtedly true that they also contain 
V or general cutaneous components also. If they contain both we 
must suppose that the communis fibres are destined for the end 
buds. CoLE (98b) treats the ramus lateralis accessorius at length, 
reviewing the literature and giving the results of anatomical and 
microscopical investigation in Gadus. He finds the nerve arising 
from the “trigeminal” ganglion, from the facial fibres of the hyo- 
mandibular trunk, from the vagus root, and from the roots of spinal 
nerves. From this it appears that at least the majority of the fibres 
in this nerve must be communis fibres since those of the facialis 
are such and those of the vagus are probably so. There is also, it 
seems from the description, a general cutaneous component from 
the V. The most probable interpretation of the facts in the present 
state of our knowledge is that the communis fibres innervate the 
end buds of the fins, while the V fibres are merely general cuta- 
11* 
