The brain of Acipenser. . 165 
culus communis of STRONG, and issues in front of and internal to 
the sense organ root”. KınasBury (97) confirms the above de- 
scriptions for Acipenser and Amia. 
For Teleosts the root has been described by MAYSER (81) as 
the “dorsal geniculate V” from the “Lobus trigemini” (= front part 
of L. vagi). KINGSBURY identifies the same root in numerous Te- 
leosts. In the cod, CoLE (’98b) describes two roots for the whole 
V—VII complex including the lateral line VII, but he has not worked 
out the several components in these roots. C. J. HERRICK (97 and 
99) describes a single root in Menidia for all the V—VII com- 
ponents except the motor VII. The sensory VII comes from the front 
end of the fasciculus communis, the whole of which goes to the 
usual sensory VII rami and the ramus lateralis accessorius. 
In Protopterus (Pınkus, 93) the separate roots of the palatinus 
and hyomandibularis arise in front of VIII, by which they are partly 
separated from the lateral line roots. STRONG (95) showed that the 
dorsal root (VII aa) arises in the tadpole from the fasciculus communis 
= lobus vagi of fishes) and forms the palatinus and mandibularis 
internus (chorda tympani) which supply the front part of the pharynx, 
including the end buds there situated. STRONG first pointed out 
that this root corresponds to the intermediary nerve of WRISBERG 
in man. 
The other nerves of the communis system are the sensory IX 
and X. These have their end-nuclei in the lobus vagi in fishes 
(fasciculus communis in Amphibia) and their fibres go to innervate 
the mucous lining of the branchial and pharyngeal cavities, the in- 
testine and other viscera, and to help form the ramus lateralis ac- 
cessorius which innervates end buds on the body and fins (ALLIS, 
STRONG, HERRICK, and others). In some cases also IX and X com- 
munis fibres probably go to end buds by way of the lateral line X 
(HERRICK, ’99). 
1) Nucleus funiculi et trigemini spinalis. 
This nucleus has not before been studied by the GoLcı method 
in fishes. MAYSER (81) describing the Cyprinoids, says that in the 
dorsal tracts of the cord the column of GOLL is apparently absent. 
The dorsal tract partly ends in the enlarged dorsal horn at the ce- 
phalic end of the cord and partly joins the spinal V. The spinal V 
contains three kinds of fibres: rather fine with or without light me- 
dullation; medium, heavily medullated; and extremely fine fibres. 
