Nachdruck verholen, 



Ueher setzungsrecht vorbehalten. 



The Origin and Development of the Wings 

 of Coleoptera. 



By 

 W. L. Tower. 



With plates 14 — 20 and 8 figures in the text. 



Contents. 

 I. Introduction. 

 II. Development of the Wings in Coleoptera. 



1. Origin of the Wing Fundament. 



a) Embryonic Development. 



b) Growth and Invagination in the Larva. 



2. Formation of the larval Wings. 



3. Significance of the Types of W^ing Development. 



4. Behavior of the Hypodermis. 



5. Tracheal System of the Wings. 



a) Larval tracheal System. 



b) Adult tracheal System. 



6. Formation of the Veins. 



7. Other Tissues in the Larval Wings. 



8. Homology of the Elytra. 



9. The prepupal Stage and Pupation. 



10. Development in the Papa. 



11. The mature Wings. 



a) Elytra. 



b) Hind Wings. 

 III. Conclusion. 



Methods. 

 Bibliography. 

 Explanation of Plates. 



I. Introduction. 



That the wings of insects are present in the larval stage was 

 first discovered by Malpighi, in 1687, who found them already well 

 developed in the mature larva of Bombyx mori. Following Malpighi, 



