238 ALUS. [Vol. XVIII. 



always found a small strand which separated from the main nerve 

 but was distributed, like the remainder of the nerve, to the rectus 

 externus. It however ran forward a considerable distance along 

 the muscle before entering it, instead of entering it at once, as the 

 larger part of the nerve did. 



7. Trigemino-Facial Complex. 



The Radix Profundi, usually considered as a part of the 

 trigemino-facial complex, arises from the brain separate and dis- 

 tinct from the other roots of the complex. It lies antero-mesial 

 to the latter roots, runs forward and outward and enters a small 

 ganglion {gp) while still inside the cranial cavity. This ganglion 

 may be connected by a nervous strand, perhaps sympathetic, with 

 the root of the nervus trigeminus. From the ganglion, which is 

 evidently the homologue of the whole or a part of the profundus 

 ganglion of Atnia, a single nerve arises, which I have called the 

 truncus ciliaris profundi. Immediately after leaving its ganglion 

 this truncus traverses the profundus foramen in the petrosal, and 

 enters the trigemino-facial chamber on the outer surface of that 

 bone. Issuing from the chamber, by its trigeminal opening, the 

 truncus separates, sooner or later, into two parts, one of .which is the 

 radix longa and the other the ciliaris longus, the manner and place 

 of separation of the two parts varying in different specimens, and 

 three strands often being found (Fig. 66), two of which together 

 form the radix longa. Either the truncus ciliaris profundi alone, 

 or both the truncus and the ciliaris longus, were always connected, 

 by a variable number of strands, with a large sympathetic ganglion 

 found intimately associated with the trigeminal nerves. Whether 

 one or more of these strands represented the portio ophthalmici 

 profundi of Amia, or not, could not be determined from dissection. 

 They seemed however, from their course and general appearance, 

 to be simply sympathetic fibers sent from the sympathetic ganglion 

 to the ciliary ganglion and nerves. If this be so the portio ophthal- 

 mici profundi of Amia must either be wholly wanting in Scomber, 

 or wholly fused with the trigeminus, those parts of the profundus 

 ganglion and radix profundi that belong to it included. For the 

 present it seems to me better to consider the nerve as wholly want- 

 ing in Scomber, as I was l&d to consider it to be in Trigla (No. 



