Nos. IAXD2.] ANATOMY OF SCOMBER SCOMBER. 251 



stated in describing the lateral canals, by a branch of the buccalis 

 facialis. 



It is thus evident that the ophthalmicus facialis of Scomber 

 must end in that branch of the united superficial nerves that goes 

 to organ No. 2 supraorbital, and that beyond that branch the fibers 

 of the trunk are wholly trigeminal. It is then the ophthalmicus 

 trigemini alone that anastomoses with the maxillaris superior on 

 the top of the snout. This anastomosis of these two nerves was 

 not found in Amia, although the two nerves of that fish approach 

 each other distally ; nor is it described by Goronowitsch or Cole in 

 either Lota or Gadns. 



In the young of Gad us Cole says (Xo. 16, p. 156) that the two 

 ophthalmic nerves, a certain distance beyond their points of origin 

 from the ganglionic complex, enter "what appears to be a rudi- 

 mentary eye muscle canal." As that part of the ganglionic com- 

 plex that lies in front of the truncus hyoideo-mandibularis is said 

 by Cole {loc. cit., p. 135) to lie partly inside and partly outside the 

 cranium, and as the ophthalmicus trigemini is given off from the 

 anterior end of the ganglion, this nerve, at least, must have already 

 issued from the cranium before entering the canal in question. 

 Both nerves are said to leave the canal, in the "region of sense 

 organ 3" supraorbital, that is, in the figure given, dorsal to the 

 anterior edge of the eyeball. What the homologue of this canal 

 may be, in either Amia or Scovihcr, I am wholly unable to tell. 



The Truncus Maxillaris Trigemini {tmt) arises from the 

 trigemino-facial ganglionic complex, either in the trigemino-facial 

 chamber or immediately beyond the trigeminal opening of that 

 chamber. It lies, at its origin, ventral to the buccalis facialis, 

 which nerve arises independently of it, from the dorsal, more or 

 less independent part of the complex. Lying close to the buccalis, 

 and usually connected with it by a short commissural branch, it 

 runs downward and forward internal to the levator arcus palatini, 

 and, near the anterior edge of that muscle, separates into its supe- 

 rior and inferior divisions. 



From the base of the truncus, or perhaps from what is still a 

 part of the ganglion, a large commissural branch is sent backward, 

 through the trigemino-facial chamber, to the truncus hyoideo- 

 mandibularis facialis. This branch often arises bv two strands 



