254 ALUS. [Vol. XVIII. 



says that a branch from this gangHon anastomoses with the third 

 nerve, and then " gives rise to the usual ciliary nerve." 



The AIaxillaris Superior Trigemini {mst) and Buccalis 

 Facialis {hf) of Scomber usually anastomose shortly after the 

 former nerve has separated from the maxillaris inferior. From 

 this anastomosis two nerve strands arise which continue downward 

 and forward under the eyeball, anastomosing again, one or more 

 times, or being connected with each other, in a variable manner, 

 by communicating branches. That there is, in these several anas- 

 tomoses, an important interchange of nerve elements is evident 

 from the ultimate distribution of the branches of each of the 

 two nerve strands. The two strands, in this suborbital part of 

 their course, lie external to, that is, posterior or ventral to, all 

 the muscles of the orbit and the nerves innervating them, and 

 dorsal to the floor of the orbit, that floor being formed either by 

 the adductor arcus palatini or by parts of the palato-quadrate 

 arch. 



The buccalis facialis gives off two branches before its first 

 anastomosis with the maxillaris superior. The first of these two 

 branches is the ramus oticus facialis {of), which arises from the 

 base of the buccalis, or from the ganglionic complex close to the 

 base of that nerve. Turning upward along the hind wall of the 

 orbit it enters the large pit in the orbital face of the postorbital 

 ossification, and issues on the top of the skull by the one or two 

 openings on the dorsal surface of that same bone, near its lateral 

 edge. It there separates into two parts. One of these parts turns 

 forward, and, entering the postfrontal bone, innervates organ lo 

 of the infraorbital canal. The other part turns backward and 

 mesially, along the external surface of the dilatator operculi, and 

 separates into two branches, both of which turn inward between 

 the dilatator and the ridge of the squamosal that bounds its dorso- 

 mesial edge, and entering the ridge innervate, one of them organ 

 II infraorbital, and the other organ 12. 



The oticus facialis oi Scomber thus corresponds to the nerve 

 identified as the oticus facialis in Amia, plus that branch of the 

 buccalis of Amia that arises close to the base of the ramus oticus, 

 almost as a part of that nerve, and that goes to organ 14 infra- 

 orbital. The otic part of the main infraorbital canal of Scomber 



